Yucca Brevifolia
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''Yucca brevifolia'' is a plant species belonging to the genus ''
Yucca ''Yucca'' is a genus of perennial plant, perennial shrubs and trees in the family (biology), family Asparagaceae, subfamily Agavoideae. Its 40–50 species are notable for their Rosette (botany), rosettes of evergreen, tough, sword-shaped Leaf, ...
''. It is tree-like in habit, which is reflected in its common names: Joshua tree, yucca palm, tree yucca, and palm tree yucca. This
monocotyledon Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one of ...
ous
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
is native to the arid
Southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, Ne ...
, specifically
California California is a U.S. state, state in the Western United States, located along the West Coast of the United States, Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the List of states and territori ...
,
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
,
Utah Utah ( , ) is a state in the Mountain West subregion of the Western United States. Utah is a landlocked U.S. state bordered to its east by Colorado, to its northeast by Wyoming, to its north by Idaho, to its south by Arizona, and to it ...
, and
Nevada Nevada ( ; ) is a U.S. state, state in the Western United States, Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. N ...
, and to northwestern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
It is confined mostly to the
Mojave Desert The Mojave Desert ( ; mov, Hayikwiir Mat'aar; es, Desierto de Mojave) is a desert in the rain shadow of the Sierra Nevada mountains in the Southwestern United States. It is named for the indigenous Mojave people. It is located primarily in ...
between elevation. It thrives in the open grasslands of Queen Valley and
Lost Horse Valley Lost Horse Valley is a valley in Joshua Tree National Park. It has been called "the centerpiece of the park" because of the quality of its stands of Joshua trees and its geological formations. It contains the Lost Horse Mine Lost Horse Mine is a ...
in
Joshua Tree National Park Joshua Tree National Park is an American national park in southeastern California, east of San Bernardino and Los Angeles and north of Palm Springs. It is named after the Joshua trees (''Yucca brevifolia'') native to the Mojave Desert. Origin ...
. Other regions with large populations of the tree can be found northeast of Kingman, Arizona in
Mohave County Mohave County is in the northwestern corner of the U.S. state of Arizona. As of the 2020 census, its population was 213,267. The county seat is Kingman, and the largest city is Lake Havasu City. It is the fifth largest county in the United St ...
; and along U.S. 93 between the towns of
Wickenburg Wickenburg is a town in Maricopa and Yavapai counties, Arizona, United States. As of the 2020 census, the population of the town was 7,474, up from 6,363 in 2010. History The Wickenburg area, along with much of the Southwest, became part of ...
and Wikieup, a route which has been designated the Joshua Tree Parkway of Arizona. The common name Joshua tree apparently comes from Christian iconography.


Taxonomy

The Joshua tree is also called ''izote de desierto'' (Spanish, "desert dagger"). It was first formally described in the botanical literature as ''Yucca brevifolia'' by
George Engelmann George Engelmann, also known as Georg Engelmann, (2 February 1809 – 4 February 1884) was a German-American botanist. He was instrumental in describing the flora of the west of North America, then very poorly known to Europeans; he was particu ...
in 1871 as part of the Geological Exploration of the 100th meridian (or "
Wheeler Survey The Wheeler Survey, carried out in 1872-1879, was one of the "Four Great Surveys" conducted by the US government after the Civil War primarily to document the geology and natural resources of the American West. Supervised by First Lieutenant (lat ...
"). The name "Joshua tree" is commonly said to have been given by a group of
Mormon Mormons are a religious and cultural group related to Mormonism, the principal branch of the Latter Day Saint movement started by Joseph Smith in upstate New York during the 1820s. After Smith's death in 1844, the movement split into several ...
settlers crossing the Mojave Desert in the mid-19th century: The tree's role in guiding them through the desert combined with its unique shape reminded them of a biblical story in which
Joshua Joshua () or Yehoshua ( ''Yəhōšuaʿ'', Tiberian: ''Yŏhōšuaʿ,'' lit. 'Yahweh is salvation') ''Yēšūaʿ''; syr, ܝܫܘܥ ܒܪ ܢܘܢ ''Yəšūʿ bar Nōn''; el, Ἰησοῦς, ar , يُوشَعُ ٱبْنُ نُونٍ '' Yūšaʿ ...
keeps his hands reached out for an extended period of time to enable the Israelites in their conquest of Canaan (). Further, the shaggy leaves may have provided the appearance of a beard. However, no direct or contemporary attestation of this origin exists, and the name Joshua tree is not recorded until after Mormon contact; moreover, the physical appearance of the Joshua tree more closely resembles a similar story told of
Moses Moses hbo, מֹשֶׁה, Mōše; also known as Moshe or Moshe Rabbeinu (Mishnaic Hebrew: מֹשֶׁה רַבֵּינוּ, ); syr, ܡܘܫܐ, Mūše; ar, موسى, Mūsā; grc, Mωϋσῆς, Mōÿsēs () is considered the most important pro ...
. Ranchers and miners who were contemporaneous with the Mormon immigrants used the trunks and branches as fencing and for fuel for ore-processing steam engines. They referred to these fallen or collapsed Joshua trees as ''tevis'' stumps. In addition to the autonymic
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
''Y. b.'' subsp. ''brevifolia'', two other subspecies have been described: ''Y. b.'' subsp. ''herbertii'' (Webber's yucca or Herbert Joshua tree) and ''Y. b.'' subsp. ''jaegeriana'' (the Jaeger Joshua tree or Jaeger's Joshua tree or pygmae yucca), though both are sometimes treated as
varieties Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
or
forms Form is the shape, visual appearance, or configuration of an object. In a wider sense, the form is the way something happens. Form also refers to: *Form (document), a document (printed or electronic) with spaces in which to write or enter data * ...
. ''Y.b.'' subsp. ''jaegeriana'' has also been treated as its own species.


Growth and development

Joshua trees are fast growers for a desert species; new seedlings may grow at an average rate of per year in their first 10 years, then only about per year. The trunk consists of thousands of small
fiber Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
s and lacks annual
growth ring Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atmo ...
s, making determining the tree's age difficult. This tree has a top-heavy branch system, but also what has been described as a "deep and extensive" root system, with roots reaching down to . If it survives the rigors of the desert, it can live for hundreds of years; some specimens survive a thousand years. The tallest trees reach about . New plants can grow from
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...
, but in some populations, new stems grow from underground
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s that spread out around the parent tree. The
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
are dark green, linear, bayonet-shaped, 15–35 cm long, and 7–15 mm broad at the base, tapering to a sharp point; they are borne in a dense spiral arrangement at the apex of the stems. The leaf margins are white and
serrate Serration is a saw-like appearance or a row of sharp or tooth-like projections. A serrated cutting edge has many small points of contact with the material being cut. By having less contact area than a smooth blade or other edge, the applied pr ...
d.
Flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
s typically appear from February to late April, in
panicle A panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a panicle are of ...
s 30–55 cm tall and 30–38 cm broad, the individual flowers erect, 4–7 cm tall, with six creamy white to green
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s. The tepals are
lanceolate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...
and are fused to the middle. The fused
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ''pistils'' ...
s are 3 cm tall and the stigma cavity is surrounded by lobes. The semi-fleshy fruit that is produced is green-brown, elliptical, and contains many flat seeds. Joshua trees usually do not branch until after they bloom (though branching may also occur if the growing tip is destroyed by the yucca-boring
weevil Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small, less than in length, and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. They b ...
), and they do not bloom every year. Like most desert plants, their blooming depends on rainfall at the proper time. They also need a winter freeze before they bloom. Once they bloom, the flowers are pollinated by the
yucca moth The Prodoxidae are a family of moths, generally small in size and nondescript in appearance. They include species of moderate pest status, such as the currant shoot borer, and others of considerable ecological and evolutionary interest, such as ...
(''
Tegeticula synthetica ''Tegeticula synthetica'' is a moth of the family Prodoxidae. It is found in the United States in the Mojave Desert in southern Nevada, south-eastern California and from south-western Utah to north-western Arizona. The habitat consists of desert ...
''), which spreads pollen while laying eggs inside the flower. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e feed on the seeds, but enough seeds remain to reproduce. The Joshua tree is also able to actively abort ovaries in which too many eggs have been produced.


Distribution and habitat

The Joshua tree is native to the southwestern United States (Arizona, California, Nevada, and Utah) and northwestern Mexico. This range mostly coincides with the geographical reach of the Mojave Desert, where it is considered one of the major
indicator species A bioindicator is any species (an indicator species) or group of species whose function, population, or status can reveal the qualitative status of the environment. The most common indicator species are animals. For example, copepods and other sma ...
for the desert. It occurs at elevations between .


Conservation status

Joshua trees are one of the species predicted to have their range reduced and shifted by
climate change In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to E ...
. Concern remains that they will be eliminated from Joshua Tree National Park, with ecological research suggesting a high probability that their populations will be reduced by 90% of their current range by the end of the 21st century, thus fundamentally transforming the ecosystem of the park. Wildfires, invasive grasses and poor migration patterns for the trees’ seeds are all additional factors in the species’ imperilment. As an example, more than a quarter—or more than 1.3 million Joshua trees—in of one of the densest Joshua tree populations in the world in
Mojave National Preserve Mojave National Preserve is a United States National Preserve located in the Mojave Desert of San Bernardino County, California, USA, between Interstate 15 in California, Interstate 15 and Interstate 40. The preserve was established October 31 ...
were killed in the Dome Fire in August 2020. Also, concern exists about the ability of the species to migrate to favorable climates due to the extinction of the giant
Shasta ground sloth ''Nothrotheriops'' is a genus of Pleistocene ground sloth found in North America, from what is now central Mexico to the southern United States. This genus of bear-sized xenarthran was related to the much larger, and far more famous '' Megatheriu ...
(''Nothrotheriops shastensis'') 13,000 years ago; ground sloth dung has been found to contain Joshua tree leaves, fruits, and seeds, suggesting that the sloths might have been key to the trees' dispersal.


Uses and cultivation

Different forms of the species are cultivated, including smaller plants native from the eastern part of the species range. These smaller plants grow 2.5 m tall and branch when about 1 m tall.
Red-shafted flicker The northern flicker or common flicker (''Colaptes auratus'') is a medium-sized bird of the woodpecker family. It is native to most of North America, parts of Central America, Cuba, and the Cayman Islands, and is one of the few woodpecker speci ...
s make nests in the branches, which are later used by other birds.