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The yak (''Bos grunniens''), also known as the Tartary ox, grunting ox, hairy cattle, or domestic yak, is a species of long-haired
domesticated Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of reso ...
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
found throughout the Himalayan region, the
Tibetan Plateau The Tibetan Plateau, also known as the Qinghai–Tibet Plateau or Qingzang Plateau, is a vast elevated plateau located at the intersection of Central Asia, Central, South Asia, South, and East Asia. Geographically, it is located to the north of H ...
,
Tajikistan Tajikistan, officially the Republic of Tajikistan, is a landlocked country in Central Asia. Dushanbe is the capital city, capital and most populous city. Tajikistan borders Afghanistan to the Afghanistan–Tajikistan border, south, Uzbekistan to ...
, the
Pamir Mountains The Pamir Mountains are a Mountain range, range of mountains between Central Asia and South Asia. They are located at a junction with other notable mountains, namely the Tian Shan, Karakoram, Kunlun Mountains, Kunlun, Hindu Kush and the Himalaya ...
, and as far north as
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
and
Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...
. It is descended from the
wild yak The wild yak (''Bos mutus'') is a large, wild bovine native to the Himalayas. It is the ancestor of the domestic yak (''Bos grunniens''). Taxonomy The ancestor of the wild and domestic yak is thought to have diverged from '' Bos primigenius' ...
(''Bos mutus'').


Etymology

The English word ''yak'' originates from the . In Tibetan and Balti it refers only to the male of the species, the female being called or in Tibetan and in Balti. In English, as in most other languages that have borrowed the word, ''yak'' is usually used for both sexes, with ''bull'' or ''cow'' referring to each sex separately.


Taxonomy

Belonging to the genus ''
Bos ''Bos'' (from Latin '' bōs'': cow, ox, bull) is a genus of bovines, which includes, among others, wild and domestic cattle. ''Bos'' is often divided into four subgenera: ''Bos'', ''Bibos'', ''Novibos'', and ''Poephagus'', but including t ...
'', yaks are related to
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
(''Bos primigenius'').
Mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
analyses to determine the evolutionary history of yaks have been inconclusive. The yak may have diverged from cattle at any point between one and five million years ago, and there is some suggestion that it may be more closely related to
bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...
than to the other members of its designated genus. Apparent close fossil relatives of the yak, such as ''
Bos baikalensis ''Bos'' (from Latin '' bōs'': cow, ox, bull) is a genus of bovines, which includes, among others, wild and domestic cattle. ''Bos'' is often divided into four subgenera: ''Bos'', ''Bibos'', ''Novibos'', and ''Poephagus'', but including the ...
'', have been found in eastern Russia, suggesting a possible route by which yak-like ancestors of the modern
American bison The American bison (''Bison bison''; : ''bison''), commonly known as the American buffalo, or simply buffalo (not to be confused with Bubalina, true buffalo), is a species of bison that is endemic species, endemic (or native) to North America. ...
could have entered the Americas. The species was originally designated as ''Bos grunniens'' ("grunting ox") by
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
in 1766. Still, this name is now generally considered to refer only to the domesticated form of the animal, with ''Bos mutus'' ("mute ox") being the preferred name for the wild species. Although some authors still consider the wild yak to be a
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
, ''Bos grunniens mutus'', the
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...
made an official ruling in 2003 permitting the use of the name ''Bos mutus'' for
wild yak The wild yak (''Bos mutus'') is a large, wild bovine native to the Himalayas. It is the ancestor of the domestic yak (''Bos grunniens''). Taxonomy The ancestor of the wild and domestic yak is thought to have diverged from '' Bos primigenius' ...
s, and this is now the more common usage. There are no recognised subspecies of yak except where the wild yak is considered a subspecies of ''Bos grunniens''.


Physical characteristics

Yaks are heavily built animals with bulky frames, sturdy legs, rounded, cloven hooves, and extremely dense, long fur hanging lower than the belly. While wild yaks are generally dark, blackish to brown in colouration, domestic yaks can be quite variable, often having rusty brown and cream patches. They have small ears and broad foreheads, with smooth horns that are generally dark in colour. In males (bulls), the horns sweep out from the sides of the head and then curve backwards; they typically range from in length. The horns of females (cows) are smaller, at in length, and have a more upright shape. Both sexes have a short neck with a pronounced hump over the shoulders, although this is larger and more visible in males. Males weigh , females weigh . Wild (feral) yaks can be substantially heavier, bulls reaching weights of up to . Depending on the breed, domestic yak males are high at the withers, while females are high at the withers. Both sexes have long, shaggy hair with a dense woolly undercoat over the chest, flanks, and thighs to insulate them from the cold. Especially in bulls, this may form a long "skirt" that can reach the ground. The tail is long and horselike rather than tufted like the tails of cattle or bison. Domesticated yaks have a wide range of coat colours, with some individuals being white, grey, brown, roan or
piebald A piebald or pied animal is one that has a pattern of unpigmented spots (white) on a pigmented background of hair, feathers or scales. Thus a piebald black and white dog is a black dog with white spots. The animal's skin under the white backg ...
. The
udder An udder is an organ formed of two or four mammary glands on the females of dairy animals and ruminants such as cattle, goats, and sheep. An udder is equivalent to the breast in primates, elephantine pachyderms and other mammals. The udder is ...
in females and the
scrotum In most terrestrial mammals, the scrotum (: scrotums or scrota; possibly from Latin ''scortum'', meaning "hide" or "skin") or scrotal sac is a part of the external male genitalia located at the base of the penis. It consists of a sac of skin ...
in males are small and hairy as protection against the cold. Females have four
teat A teat is the projection from the mammary glands of mammals from which milk flows or is ejected for the purpose of feeding young. In many mammals, the teat projects from the udder. The number of teats varies by mammalian species and often corr ...
s. Yaks are not known to produce the characteristic lowing (mooing) sound of cattle, but both wild and domestic yaks grunt and squeak, which inspired the scientific name of the domestic yak variant, ''Bos grunniens'' (grunting bull).
Nikolay Przhevalsky Nikolay Mikhaylovich Przhevalsky (or Prjevalsky;; , . – ) was a Russian geographer and a renowned explorer of Central and East Asia. Although he never reached his ultimate goal, the city of Lhasa in Tibet, he still travelled through regio ...
named the wild variant ''Bos mutus'' (silent bull), believing that it did not make a sound at all, but it does.


Physiology

Yak physiology is well adapted to high altitudes, having larger lungs and heart than cattle found at lower altitudes, as well as greater capacity for transporting oxygen through their blood, due to the persistence of
foetal haemoglobin Fetal hemoglobin, or foetal haemoglobin (also hemoglobin F, HbF, or α2γ2) is the main oxygen carrier protein in the human fetus. HemoglobinF is found in fetal red blood cells, and is involved in transporting oxygen from the mother's bloodstream ...
throughout life. Conversely, yaks have trouble thriving at lower altitudes, and are prone to suffering from heat exhaustion above about . Further adaptations to the cold include a thick layer of
subcutaneous fat The subcutaneous tissue (), also called the hypodermis, hypoderm (), subcutis, or superficial fascia, is the lowermost layer of the integumentary system in vertebrates. The types of cells found in the layer are fibroblasts, adipose cells, and ...
and an almost complete lack of functional
sweat glands Sweat glands, also known as sudoriferous or sudoriparous glands, , are small tubular structures of the skin that produce sweat. Sweat glands are a type of exocrine gland, which are glands that produce and secrete substances onto an epithelial sur ...
. Compared with domestic cattle, the
rumen The rumen, also known as a paunch, is the largest stomach compartment in ruminants. The rumen and the reticulum make up the reticulorumen in ruminant animals. The diverse microbial communities in the rumen allows it to serve as the primary si ...
of yaks is unusually large, relative to the
omasum The omasum, also known as the green, the fardel, the manyplies and the psalterium, is the third compartment of the stomach in ruminants. The omasum comes after the rumen and reticulum and before the abomasum. Different ruminants have different oma ...
. This likely allows them to consume greater quantities of low-quality food at a time, and to ferment it longer to extract more nutrients. Yak consume the equivalent of 1% of their body weight daily while cattle require 3% to maintain condition. They are grazing herbivores, with their wild ancestors feeding primarily on
grass Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family (biology), family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and spe ...
and
sedges The Cyperaceae () are a family of graminoid (grass-like), monocotyledonous flowering plants known as sedges. The family is large; botanists have described some 5,500 known species in about 90 generathe largest being the "true sedges" (genu ...
, with some herbs and dwarf shrubs.


Reproduction and life history

Yaks mate in the summer, typically between July and September, depending on the local environment. For the remainder of the year, many bulls wander in small bachelor groups away from the large herds. Still, as the rut approaches, they become aggressive and regularly fight with each other to establish dominance. In addition to non-violent threat displays, bellowing, and scraping the ground with their horns, bull yaks compete more directly, repeatedly charging at each other with heads lowered or sparring with their horns. Like
bison A bison (: bison) is a large bovine in the genus ''Bison'' (from Greek, meaning 'wild ox') within the tribe Bovini. Two extant taxon, extant and numerous extinction, extinct species are recognised. Of the two surviving species, the American ...
, but unlike cattle, males wallow in dry soil during the rut, often while scent-marking with urine or dung. Females enter
oestrus The estrous cycle (, originally ) is a set of recurring physiological changes induced by reproductive hormones in females of mammalian subclass Theria. Estrous cycles start after sexual maturity in females and are interrupted by anestrous phas ...
up to four times a year, and females are receptive only for a few hours in each cycle.
Gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...
lasts between 257 and 270 days, so that the young are born between May and June, and results in the birth of a single calf. The cow finds a secluded spot to give birth, but the calf can walk within about ten minutes of birth, and the pair soon rejoin the herd. Females of both the wild and domestic forms typically give birth only once every other year, although more frequent births are possible if the food supply is good. Calves are
weaned Weaning is the process of gradually introducing an infant human or other mammal to what will be its adult diet while withdrawing the supply of its mother's milk. In the UK, weaning primarily refers to the introduction of solid foods at 6 mont ...
at one year and become independent shortly thereafter. Wild calves are initially brown in color and only later develop darker adult hair. Females generally give birth for the first time at three or four years of age, and reach their peak reproductive fitness at around six years. Yaks may live for more than twenty years in domestication or captivity, although it is likely that this may be somewhat shorter in the wild.


Husbandry

For thousands of years, domesticated yaks have been kept in Mongolia and Tibet, primarily for their milk,
fibre Fiber (spelled fibre in British English; from ) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorp ...
(wool), and meat, and as
beasts of burden A working animal is an animal, usually domesticated, that is kept by humans and trained to perform tasks. Some are used for their physical strength (e.g. oxen and draft horses) or for transportation (e.g. riding horses and camels), while oth ...
. Their dried droppings are an important fuel, used all over Tibet, and are often the only fuel available on the high, treeless
Tibetan Plateau The Tibetan Plateau, also known as the Qinghai–Tibet Plateau or Qingzang Plateau, is a vast elevated plateau located at the intersection of Central Asia, Central, South Asia, South, and East Asia. Geographically, it is located to the north of H ...
. Yaks transport goods across mountain passes for local farmers and traders and are an attraction for climbing and trekking expeditions: "Only one thing makes it hard to use yaks for long journeys in barren regions. They will not eat
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
, which could be carried on the journey. They will starve unless they can be brought to a place where there is grass." They also are used to draw
plough A plough or ( US) plow (both pronounced ) is a farm tool for loosening or turning the soil before sowing seed or planting. Ploughs were traditionally drawn by oxen and horses but modern ploughs are drawn by tractors. A plough may have a wooden ...
s. Yaks' milk is often processed to a cheese called ''
chhurpi Chhurpi (), otherwise known as ''durkha'' and ''chogo/chugo'', is a traditional cheese consumed in Nepal, Bhutan and parts of Northeastern India. The two varieties of chhurpi are a soft variety (consumed usually as a side dish with rice) and a ve ...
'' in Tibetan and Nepali languages, and '' byaslag'' in Mongolia. Butter made from yaks' milk is an ingredient of the
butter tea Butter tea, also known as ''Bho jha'' (, "Tibetan tea"), ''cha süma'' (, "churned tea", Mandarin Chinese: ''sūyóu chá'' ( 酥 油 茶), ''su ja'' (, "churned tea") in Dzongkha, ''Cha Su-kan'' or "gur gur cha" in the Ladakhi language and Su C ...
that Tibetans consume in large quantities, and is also used in lamps and made into butter sculptures used in religious festivities.


Outside the Himalayas

Small numbers of herds can be found in the United States, Canada, New Zealand, and some parts of Europe. Yaks have generated interest outside the Himalayas as a commercial crop and by cattle breeders. The main interest of North American yak breeders is lean meat production by hybridizing with other cattle, followed by
wool Wool is the textile fiber obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have some properties similar to animal w ...
production.


Research

The
Indian government The Government of India (ISO: Bhārata Sarakāra, legally the Union Government or Union of India or the Central Government) is the national authority of the Republic of India, located in South Asia, consisting of 36 states and union territor ...
established a dedicated centre for research into yak husbandry, the ICAR-National Research Centre on Yak, in 1989. It is located at
Dirang Dirang H.Q. (simply Dirang) is a village in the district of West Kameng in the Indian state of Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-Ea ...
, Arunachal Pradesh, and maintains a yak farm in the Nyukmadung area at an altitude of above MSL.


Breeding and hybridization

In
Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, and
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
, domestic cattle are crossbred with yaks. This gives rise to the infertile male dzo མཛོ། as well as fertile females known as or མཛོ་མོ།, which may be crossed again with cattle. The Dwarf Lulu breed, "the only ''Bos primigenius taurus'' type of cattle in Nepal" has been tested for DNA markers and found to be a mixture of both taurine and
zebu The zebu (; ''Bos indicus''), also known as indicine cattle and humped cattle, is a species or subspecies of Bos taurus, domestic cattle originating in South Asia. Zebu, like many Sanga cattle breeds, differs from taurine cattle by a fatty hump ...
types of cattle (''B. p. taurus'' and ''B. p. indicus'') with yak. According to the
International Veterinary Information Service International is an adjective (also used as a noun) meaning "between nations". International may also refer to: Music Albums * ''International'' (Kevin Michael album), 2011 * ''International'' (New Order album), 2002 * ''International'' (The T ...
, the low productivity of second-generation cattle–yak crosses makes them suitable only as meat animals. Crosses between yaks and domestic cattle (''Bos primigenius taurus'') have been recorded in Chinese literature for at least 2,000 years. Successful crosses have also been recorded between yak and American bison,Zhang, R.C. (14 December 2000)
"Interspecies Hybridization between Yak, ''Bos taurus'' and ''Bos indicus'' and Reproduction of the Hybrids"
In: ''Recent Advances in Yak Reproduction'', Zhao, X.X.; Zhang, R.C. (eds.). International Veterinary Information Service.
gaur The gaur (''Bos gaurus''; ) is a large bovine native to the Indian Subcontinent and Southeast Asia, and has been listed as Vulnerable species, Vulnerable on the IUCN Red List since 1986. The global population was estimated at a maximum of 21,000 ...
, and
banteng The banteng (''Bos javanicus''; ), also known as tembadau, is a species of wild Bovinae, bovine found in Southeast Asia. The head-and-body length is between . Wild banteng are typically larger and heavier than their Bali cattle, domesticated ...
, generally with similar results to those produced with domestic cattle.


Domestication

Jacques et al. (2021) show that most elaborate yak-related terminologies are found within Tibetic and
Gyalrongic The Gyalrongic languages (also known as Rgyalrongic or Jiarongic) constitute a branch of the Qiangic languages of Sino-Tibetan languages, Sino-Tibetan, but some propose that it may be part of a larger Rung languages group and do not consider it t ...
languages. Both branches also have native terms for yak-cattle hybrids, suggesting that Tibetic and Gyalrongic speakers may have independently cross-bred yaks and cattle, predating the proto-Gyalrongic split (3221 169–4319 BP) from Tibeto-Gyalrongic. The oldest dated physical evidence of yak domestication is from 2,500 years BP.


Customs

In parts of Tibet and Karakorum,
yak racing Yak racing () is a spectator sport held at many traditional festivals of Tibet, Kazakhstan, Kyrgyzstan, Gilgit-Baltistan, Ladakh and Mongolia, in gatherings which integrate popular dances and Song, songs with traditional physical games. For Tibeta ...
is a form of entertainment at traditional festivals and an important part of their culture. More recently, sports involving domesticated yaks, such as
yak skiing Yak skiing is a sport practiced in the Indian hill resort of Manali, Himachal Pradesh, as a tourist attraction. The sport involves a skier waiting at the bottom of a slope and a yak at the top of the hill; yak and skier are connected by means of ...
or
yak polo Yak polo (or ''sarlagan polo'') is a Mongolian variant of the sport polo. It is played on yaks instead of on horseback. Originally invented in the early 2000s as a tourist attraction, the sport is as of 2006 reported to have attracted a modest ...
, are being marketed as tourist attractions in South Asian countries, including in
Gilgit-Baltistan Gilgit-Baltistan (; ), formerly known as the Northern Areas, is a region administered by Pakistan as an administrative units of Pakistan, administrative territory and consists of the northern portion of the larger Kashmir region, which has b ...
, Pakistan. In Nepal, an annual festival is held to drink the fresh blood of yak, and it is believed that it cures various diseases such as gastritis,
jaundice Jaundice, also known as icterus, is a yellowish or, less frequently, greenish pigmentation of the skin and sclera due to high bilirubin levels. Jaundice in adults is typically a sign indicating the presence of underlying diseases involving ...
, and body strain. The fresh blood is extracted from the neck of a yak without killing it. The cut is healed after the ceremony is over. The ritual is believed to be originated in Tibet and
Mustang The mustang is a free-roaming horse of the Western United States, descended from horses brought to the Americas by the Spanish conquistadors. Mustangs are often referred to as wild horses, but because they are descended from once-domesticate ...
. Traditionally, yaks are used to transport deceased people to
charnel ground A charnel ground (Sanskrit: श्मशान; IAST: śmaśāna; Tibetan pronunciation: durtrö; ) is an above-ground site for the putrefaction of bodies, generally human, where formerly living tissue is left to decompose uncovered. Although it ...
s for
sky burial Sky burial (, "bird-scattered") is a funeral practice in which a human corpse is placed on a mountaintop to decompose while exposed to the mahābhūta, elements or to be eaten by Scavenger, scavenging animals, especially vultures, bears and j ...
s in Tibet, with Tibetan practice holding that the yaks be set free upon the conclusion of the task.


Gallery

File:Yaks in Manali.jpg, Yaks in
Manali, Himachal Pradesh Manali () is a resort town, near Kullu town in Kullu district in the Indian state of Himachal Pradesh. It is situated in the northern end of the Kullu Valley, formed by the Beas River. The town is located in the Kullu district, approximately ...
, India, saddled for riding File:LitangMonastery.jpg, Train of pack yaks at Litang monastery in
Garzê Tibetan Autonomous Prefecture Garzê Tibetan Autonomous Prefecture, often shortened to Ganzi Prefecture, is an autonomous prefecture in the western arm of Sichuan province, China bordering Yunnan to the south, the Tibet Autonomous Region to the west, and Gansu to the north ...
, Sichuan, China File:Yaks still provide the best way to plow fields in Tibet.jpg, Yaks plowing fields in Tibet File:YakRace.jpg, Yaks in
Gilgit-Baltistan Gilgit-Baltistan (; ), formerly known as the Northern Areas, is a region administered by Pakistan as an administrative units of Pakistan, administrative territory and consists of the northern portion of the larger Kashmir region, which has b ...
, Pakistan File:四川 阿坝 白牦牛 - panoramio.jpg, Domestic yak in
Mao County Mao County or Maoxian ( zh, s=茂县; ; Qiangic languages, Qiang: Shgvunyi) is a counties of China, county in Ngawa Tibetan and Qiang Autonomous Prefecture, Ngawa Prefecture, Sichuan, Sichuan Province, China. It has an area of 3,903 and a popul ...
, China File:四姑娘山景区 Mount Siguniang Scenic Area 17.jpg, A yak at Mount Siguniang Scenic Area, Sichuan, China


See also

*
Yakalo The yakalo is a cross of the yak (''Bos grunniens'') and the American bison (''Bison bison'', known as a buffalo in North America). It was produced by hybridisation experiments in the 1920s, when crosses were made between yak bulls and both pur ...
* Yak dance


References


External links

* *
International Yak Association (IYAK)European Yak Association (EYAK)Yak Genome Database
{{Taxonbar, from=Q19829410 Livestock Domesticated animals Pack animals Mammals of Central Asia Mammals of East Asia Mammals of Siberia Mammals of South Asia Mammals of Tibet Fauna of the Himalayas Mammals described in 1766 Taxa named by Carl Linnaeus Bos