Vicia Orobus
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''Vicia orobus'' is a species of leguminous plant in the genus '' Vicia'', known as wood bitter-vetch. It is found in Atlantic areas of Europe, especially in the rocky edges of seasonally-grazed fields. It grows up to tall, and has no
tendril In botany, a tendril is a specialized Plant stem, stem, leaf or Petiole (botany), petiole with a thread-like shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There ar ...
s at the ends of its pinnate leaves. Its
flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s are white with purple veins, and are borne in groups of 6 or more.


Description

''Vicia orobus'' is a
perennial plant In horticulture, the term perennial (''wikt:per-#Prefix, per-'' + ''wikt:-ennial#Suffix, -ennial'', "through the year") is used to differentiate a plant from shorter-lived annual plant, annuals and biennial plant, biennials. It has thus been d ...
, growing up to tall. Its leaves are paripinnate, with 6–15 pairs of leaflets on each leaf. The flowers are borne in groups of 6–20. Each individual flower is long, and is white with purple veins. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is a 4–5-seeded pod around long. ''V. orobus'' can be distinguished from other species of ''Vicia'' occurring in the British Isles by a number of characters. It is one of three species to lack
tendril In botany, a tendril is a specialized Plant stem, stem, leaf or Petiole (botany), petiole with a thread-like shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There ar ...
s (the others being '' V. lathyroides'' and '' V. faba''), with the leaves terminating instead in a short point. It differs from the other two species without tendrils in being perennial, having more than 6 flowers in each
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
, having peduncles more than long, and having more than 5 pairs of leaflets in each pinnate leaf. The lack of tendrils is thought to represent a primitive state within the genus ''Vicia''.


Distribution

''Vicia orobus'' has an Atlantic distribution, occurring from northern
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
to
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of the Kingdom of ...
, at altitudes of . In
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
, ''V. orobus'' has a westerly distribution, and is extinct in
South East England South East England is one of the nine official regions of England, regions of England that are in the ITL 1 statistical regions of England, top level category for Statistics, statistical purposes. It consists of the nine counties of england, ...
. Much of the global population is found in the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
, where it is largely restricted to altitudes of , except in the far north (north of Lochinver), where it can be found down to
sea level Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...
. Its stronghold is in central and north-western
Wales Wales ( ) is a Countries of the United Kingdom, country that is part of the United Kingdom. It is bordered by the Irish Sea to the north and west, England to the England–Wales border, east, the Bristol Channel to the south, and the Celtic ...
. In
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
, ''V. orobus'' is declining in numbers, and has been found recently at fewer than a dozen sites, most of which hold very small populations. It is therefore accorded legal protection in the Republic of Ireland, under the ''Flora (Protection) Order 1999''. In continental Europe, ''V. orobus'' is replaced to the east by '' V. cassubica'', with the division between the two species running through central France. It is present as a native species in
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
, where it is the only truly Atlantic species.


Habitat

''Vicia orobus'' is found in a variety of sites, including the edges of woods, on heaths, in
meadow A meadow ( ) is an open habitat or field, vegetated by grasses, herbs, and other non- woody plants. Trees or shrubs may sparsely populate meadows, as long as they maintain an open character. Meadows can occur naturally under favourable con ...
s or in rocky places over
limestone Limestone is a type of carbonate rock, carbonate sedimentary rock which is the main source of the material Lime (material), lime. It is composed mostly of the minerals calcite and aragonite, which are different Polymorphism (materials science) ...
. In Great Britain, it is frequently found at the edges of fields where
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
are grazed in winter, but a hay crop is grown in summer. It is threatened by both overgrazing and undergrazing, as well as other activities, such as grassland improvement and
land reclamation Land reclamation, often known as reclamation, and also known as land fill (not to be confused with a waste landfill), is the process of creating new Terrestrial ecoregion, land from oceans, list of seas, seas, Stream bed, riverbeds or lake ...
. ''V. orobus'' flowers from May to July and is pollinated by bees. Many species are associated with ''V. orobus'' in Great Britain, including '' Alchemilla glabra'', '' Carex pallescens'', '' Genista anglica'', '' Pseudorchis albida'', '' Rhinanthus minor'', '' Stachys officinalis'' and ''
Viola lutea The viola ( , () ) is a string instrument of the violin family, and is usually bow (music), bowed when played. Violas are slightly larger than violins, and have a lower and deeper sound. Since the 18th century, it has been the middle or alto v ...
''. At some sites in Ireland, ''V. orobus'' is associated with species such as '' Dryas octopetala'', '' Sesleria albicans'' and '' Geranium sanguineum'', which are characteristic of the flora of the Burren, although ''V. orobus'' does not occur in the Burren, or in floristically similar regions nearby.


Taxonomy

''Vicia orobus'' was first described by
Augustin Pyramus de Candolle Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss people, Swiss botany, botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple ...
in the 1815 work '. It is classified in the section ''Vicia'' sect. ''Cassubicae'', alongside species such as '' V. cassubica'', '' V. megalotropis'' and its likely sister species, '' V. sparsiflora''.


References


External links

* {{Taxonbar, from=Q159825 orobus Flora of Europe Plants described in 1815