''Vaccinium membranaceum'' is a species of
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
in the heath family
Ericaceae
The Ericaceae () are a Family (biology), family of flowering plants, commonly known as the heath or heather family, found most commonly in acidic and infertile growing conditions. The family is large, with about 4,250 known species spread acros ...
, known by the common names thinleaf huckleberry, tall huckleberry, big huckleberry, mountain huckleberry, square-twig blueberry, and ambiguously as "
black huckleberry".
Description
''Vaccinium membranaceum'' is an erect
shrub
A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
growing up to in maximum height.
The new twigs are yellow-green and somewhat angled. The
deciduous
In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
leaves are alternately arranged. The very thin to membranous, oval leaf blades are up to long. The edges are serrated,
with each tiny tooth tipped with a glandular hair.
Solitary flowers occur in the leaf axils. Each is around long,
urn-shaped to cylindrical, and pale pink to waxy bronze in color.
Cytology is 2n = 48.
They are
pollinated by
bees.
[US Forest Service Fire Ecology](_blank)
/ref> The mature fruit ranges from red through bluish-purple to a dark, almost black berry
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
about 1 cm wide. Each fruit contains an average of 47 tiny seeds.
;Reproduction
The plant rarely reproduces via seed, rather, it usually spreads by cloning
Cloning is the process of producing individual organisms with identical genomes, either by natural or artificial means. In nature, some organisms produce clones through asexual reproduction; this reproduction of an organism by itself without ...
itself from its rhizome
In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
or shoots.[ The seeds do ]germinate
Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an flowering plant, angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the sp ...
if dispersed by animals, however, as evidenced by populations of the plant growing on the recovering section of Mount St. Helens. Other than the study by Yang et al. (2008), reports of ''V. membranaceum'' sprouting from seed are quite rare with other scientists who have studied this species reporting only 6 seedlings observed during 18 years in the field.
Distribution and habitat
''Vaccinium membranaceum'' is native to western North America, with a range extending in the northern from southern Alaska
Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
, Yukon
Yukon () is a Provinces and territories of Canada, territory of Canada, bordering British Columbia to the south, the Northwest Territories to the east, the Beaufort Sea to the north, and the U.S. state of Alaska to the west. It is Canada’s we ...
, and Northwest Territories
The Northwest Territories is a federal Provinces and territories of Canada, territory of Canada. At a land area of approximately and a 2021 census population of 41,070, it is the second-largest and the most populous of Provinces and territorie ...
south as far as Utah
Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
and the northern mountains of California
California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. It can be found from the mountains next to the Pacific Ocean
The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...
in the west, to the Rocky Mountains
The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
and Black Hills in the east.
Isolated populations of this species have been found in Arizona
Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
, North Dakota
North Dakota ( ) is a U.S. state in the Upper Midwest, named after the indigenous Dakota people, Dakota and Sioux peoples. It is bordered by the Canadian provinces of Saskatchewan and Manitoba to the north and by the U.S. states of Minneso ...
, Minnesota
Minnesota ( ) is a U.S. state, state in the Upper Midwestern region of the United States. It is bordered by the Canadian provinces of Manitoba and Ontario to the north and east and by the U.S. states of Wisconsin to the east, Iowa to the so ...
, the Upper Peninsula of Michigan
The Upper Peninsula of Michigan—also known as Upper Michigan or colloquially the U.P. or Yoop—is the northern and more elevated of the two major landmasses that make up the U.S. state of Michigan; it is separated from the Lower Peninsula of ...
, and Ontario
Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...
.
''Vaccinium membranaceum'' grows at higher elevations in subalpine and alpine environments. It occurs in both pine
A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae.
''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
and spruce
A spruce is a tree of the genus ''Picea'' ( ), a genus of about 40 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal ecosystem, boreal (taiga) regions of the Northern hemisphere. ''Picea'' ...
dominated forests and in open meadow ecosystems. In forests ''V. membranaceum'' often dominates the forest understory
In forestry and ecology, understory (American English), or understorey (English in the Commonwealth of Nations, Commonwealth English), also known as underbrush or undergrowth, includes plant life growing beneath the Canopy (biology), forest ca ...
during early to mid stages of succession. ''Vaccinium membranceum'' is fire adapted. The leaves and stems of the huckleberry are resistant to low-intensity fires, and if burned away they will resprout vigorously from rhizomes buried under the soil.
Ecology
The plant also provides a key food source for black
Black is a color that results from the absence or complete absorption of visible light. It is an achromatic color, without chroma, like white and grey. It is often used symbolically or figuratively to represent darkness.Eva Heller, ''P ...
and grizzly bear
The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America.
In addition to the mainland grizzly (''Ursus arctos horr ...
s, which eat the leaves, stems, roots, and fruit. Elk, moose
The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
, and white-tailed deer
The white-tailed deer (''Odocoileus virginianus''), also known Common name, commonly as the whitetail and the Virginia deer, is a medium-sized species of deer native to North America, North, Central America, Central and South America. It is the ...
also browse the plant. The thickets provide cover for many species of small animals.
Uses
The berry is edible. ''Vaccinium membranaceum'' is the species that is the most commonly collected of all of the wild western huckleberries, and it has great commercial importance.[Flora of North America, ''Vaccinium membranaceum'' Douglas ex Torrey, 1874. Thinleaf huckleberry ](_blank)
/ref> In a good year ''V. membranaceum'' shrubs provide a lot of fruit from late summer to early fall.
Native Americans
People have been eating the fruit of this species for thousands of years. It was and continues to be widely used for food by Native Americans.[ The Kutenai called the black huckleberry shawíash ( Ktunaxa: ǂawiyaǂ). ]Alaska Natives
Alaska Natives (also known as Native Alaskans, Alaskan Indians, or Indigenous Alaskans) are the Indigenous peoples of Alaska that encompass a diverse arena of cultural and linguistic groups, including the Iñupiat, Yupik, Aleut, Eyak, Tli ...
consumed it in bread and pies as a source of vitamin C
Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...
, the Coeur d'Alene people ate the fruit fresh, dried, mashed, cooked, and added it to soup or froze it for later use, and many other groups relished it and stored it frozen, dried, pressed into cakes, or canned for winter use.
Some Native American groups lit carefully planned controlled burns in wild huckleberry patches to promote fruit production by eliminating competing plants and by stimulating the huckleberry to sprout and spread. Native American groups throughout the Pacific Northwest
The Pacific Northwest (PNW; ) is a geographic region in Western North America bounded by its coastal waters of the Pacific Ocean to the west and, loosely, by the Rocky Mountains to the east. Though no official boundary exists, the most common ...
still utilize this plant as an important cultural food and are active in its management.
In culture
The amount of fruit produced by these shrubs is legendary, with stories being told of mountain sides turned purple by all of the fruit, or shrubs being weighed to the ground by large, and abundant berries.
The huckleberry is the official state fruit of Idaho
Idaho ( ) is a landlocked U.S. state, state in the Pacific Northwest and Mountain states, Mountain West subregions of the Western United States. It borders Montana and Wyoming to the east, Nevada and Utah to the south, and Washington (state), ...
, with this particular species assumed to be the huckleberry in question.Netstate: Idaho State Fruit
/ref>
See also
*'' Gaylussacia baccata'' — which shares the common name "black huckleberry"
References
External links
Jepson Manual Treatment of ''Vaccinium membranaceum''
United States Department of Agriculture Plants Profile for ''Vaccinium membranaceum''
Washington Burke Museum, University of Washington
''Vaccinium membranaceum'' — Calphotos Photo gallery, University of California
photo of herbarium specimen at Missouri Botanical Garden, collected in Oregon in 2006
{{Authority control
membranaceum
Berries
Flora of Western Canada
Plants described in 1874
Plants used in Native American cuisine
Flora of the Northwestern United States
Flora of the Southwestern United States
Flora of South Dakota
Flora of Michigan
Flora of Ontario
Flora without expected TNC conservation status