Trielidini is a small tribe of

scoliid wasp The Scoliidae, the scoliid wasps, are a family of wasps comprising about 560 species worldwide. They tend to be black, often marked with yellow or orange, and their wing tips are distinctively corrugated. Males are more slender and elongated than ...

Description and identification

Members of Trielidini are distinguished from other members of Scoliidae by the presence of both two recurrent veins and three submarginal cells.

Taxonomy and phylogeny

This tribe was first published as Trielini by Betrem in 1965 as a tribe of Campsomerinae but was not described and was thus rendered ''

nomen nudum In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...

'' under

ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...

. Betrem published the first description under the emended name Trielidini in 1972. This tribe comprised genera with three submarginal cells as distinguished from his Campsomerini with two submarginal cells. The Campsomerinae as treated by Betrem were further distinguished from the Scoliinae by the presence of a second recurrent vein. With the discovery of '' Proscolia'' by Rasnitsyn in 1977 and establishment of the subfamily Proscoliinae, Campsomerinae was demoted to tribal status as Campsomerini, and Trielidini was treated as a symonym of Campsomerini. In his 1996 revision of scoliid higher taxonomy, Argaman established a separate subfamily, Colpinae, for Betrem's Trielidini and included tribes: Dasyscoliini, Curtaurgini, Heterelini, Colpini, and Trielidini. Argaman's subfamily and tribes, however, were established without phylogenetic analysis and were simply treated as Campsomerini in Osten's checklist of Scoliidae in 2005. A 2022 revision of North American scoliids by Kimsey ''et al.'' found through phylogenetic analysis that Campsomerini was no longer

monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...

with the inclusion of ''Colpa'' and instead found the genus was more closely allied with Scoliini, either as a member of Scoliini or as a separate tribe, suggested by the authors as Colpini. The authors opted not to establish the tribe at the time as their work did not include the other allied genera, ''Dasyscolia'' and ''Guigliana''. Mitochondrial phylogenetics conducted by Liu, van Achterberg, and Chen in 2024 validated the suspicions of Kimsey ''et al.'' and reinstated Trielidini for the genera ''Colpa'' and ''Guigliana'', without including ''Dasyscolia''. They used the name Trielidini instead of Colpini to follow ICZN guidelines.

Genera

There are two genera in the tribe Trielidini: *''

Colpa ''Colpa'' is a genus of scoliid wasp. Taxonomy and phylogeny Despite historical treatment under the tribe Campsomerini, phylogenetic analysis revealed the genus to instead be sister to the tribe Scoliini. As a result, it was excluded from Camp ...

'' Dufour, 1841 *'' Guigliana'' Betrem, 1967

References

Scoliidae Parasitic wasps Hymenoptera tribes {{Apocrita-stub