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The takin (''Budorcas taxicolor''; ), also called cattle chamois or gnu goat, is a large species of
ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...
of the subfamily
Caprinae The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine. Prominent members include sheep and g ...
found in the
eastern Himalayas ] The Eastern Himalayas extend from eastern Nepal across Northeast India, Bhutan, the Tibet Autonomous Region to Yunnan in China and northern Myanmar. The climate of this region is influenced by the monsoon of South Asia from June to September. I ...
. It includes four
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
: the Mishmi takin (''B. t. taxicolor''), the golden takin (''B. t. bedfordi''), the Tibetan (or Sichuan) takin (''B. t. tibetana''), and the Bhutan takin (''B. t. whitei''). Whilst the takin has in the past been placed together with the
muskox The muskox (''Ovibos moschatus'') is a hoofed mammal of the family Bovidae. Native to the Arctic, it is noted for its thick coat and for the strong odor emitted by males during the seasonal rut, from which its name derives. This musky odor ha ...
in the tribe Ovibovini, more recent
mitochondrial A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used ...
research shows a closer relationship to ''
Ovis ''Ovis'' is a genus of mammals, part of the Caprinae subfamily of the ruminant family (biology), family Bovidae. Its seven highly sociable species are known as sheep or ovines. Domestic sheep are members of the genus, and are thought to be des ...
'' (sheep). Its physical similarity to the muskox is therefore an example of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
. The takin is the national animal of
Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
.


Etymology

The specific name ''taxicolor'' comes from and referring to badger-like coloration.


Appearance

The takin rivals the muskox as the largest and stockiest of the subfamily
Caprinae The subfamily Caprinae, also sometimes referred to as the tribe Caprini, is part of the ruminant family Bovidae, and consists of mostly medium-sized bovids. A member of this subfamily is called a caprine. Prominent members include sheep and g ...
, which includes
goat The goat or domestic goat (''Capra hircus'') is a species of Caprinae, goat-antelope that is mostly kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the ...
s,
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
, and similar species. Its short legs are supported by large, two-toed hooves, which each have a highly developed spur. It has a stocky body and a deep chest. Its large head is distinctive by its long, arched nose and stout horns, which are ridged at the base. Horns are present in both sexes, and run parallel to the skull before turning upwards to a short point; they are about long, but can grow up to . Its long, shaggy coat is light in color with a dark stripe along the back, and males (bulls) also have dark faces. Four subspecies of takin are currently recognised, and these tend to show a variation in coat colour. Their thick
wool Wool is the textile fiber obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have some properties similar to animal w ...
often turns black in colour on their undersides and legs. Their overall coloration ranges from dark blackish to reddish-brown suffused with grayish-yellow in the eastern Himalayas to lighter yellow-gray in the
Sichuan Province Sichuan is a Provinces of China, province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capita ...
to mostly golden or (rarely) creamy-white with fewer black hairs in the
Shaanxi Province Shaanxi is a province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to the west. Shaanxi ...
. The legend of the ' golden fleece' sought by
Jason Jason ( ; ) was an ancient Greek mythological hero and leader of the Argonauts, whose quest for the Golden Fleece is featured in Greek literature. He was the son of Aeson, the rightful king of Iolcos. He was married to the sorceress Med ...
and the
Argonauts The Argonauts ( ; ) were a band of heroes in Greek mythology, who in the years before the Trojan War (around 1300 BC) accompanied Jason to Colchis in his quest to find the Golden Fleece. Their name comes from their ship, ''Argo'', named after it ...
may have been inspired by the lustrous coat of the golden takin (''B. t. bedfordi''). Hair length can range from , on the flanks of the body in summer, up to on the underside of the head in winter. In height, takin stand at the shoulder, but measure a relatively short in head-and-body length, with the tail adding only an additional . Measurements of weights vary, but according to most reports, the males are slightly larger, weighing against in females. Sources including Betham (1908) report that females are larger, with the largest captive takin known to the author, at , having been female. Takin can weigh up to or in some cases. Instead of relying on localized scent glands, the takin secretes an oily, strong-smelling substance over its whole body, enabling it to mark objects such as trees.Macdonald, D. (2001) ''The New Encyclopedia of Mammals''. Oxford University Press, Oxford. via arkive.org A prominent nose with a swollen appearance caused biologist George Schaller to liken the takin to a "bee-stung
moose The moose (: 'moose'; used in North America) or elk (: 'elk' or 'elks'; used in Eurasia) (''Alces alces'') is the world's tallest, largest and heaviest extant species of deer and the only species in the genus ''Alces''. It is also the tal ...
." Features reminiscent of familiar domesticated species have earned takins such nicknames as "cattle chamois" and "gnu goat."


Distribution and habitat

Takin are found from forested valleys to rocky, grass-covered alpine zones, at altitudes between above sea level. The Mishmi takin occurs in eastern
Arunachal Pradesh Arunachal Pradesh (; ) is a States and union territories of India, state in northeast India. It was formed from the North-East Frontier Agency (NEFA) region, and India declared it as a state on 20 February 1987. Itanagar is its capital and la ...
, while the Bhutan takin is in western Arunachal Pradesh and Bhutan. Dihang-Dibang Biosphere Reserve in Arunachal Pradesh, India is a stronghold of both Mishmi, Upper Siang (Kopu) and Bhutan takins.


Behaviour and ecology

Takin are found in small family groups of around 20 individuals, although older males may lead more solitary existences. In the summer, herds of up to 300 individuals gather high on the mountain slopes. Groups often appear to occur in largest numbers when favorable feeding sites, salt licks, or hot springs are located. Mating takes place in July and August. Adult males compete for dominance by sparring head-to-head with opponents, and both sexes appear to use the scent of their own urine to indicate dominance. A single young is born after a gestation period of around eight months. Takin migrate from the upper pasture to lower, more forested areas in winter and favor sunny spots upon sunrise.''Animal Diversity Web'' (November, 2002
"Budorcas taxicolor"
(University of Michigan Museum of Zoology) via arkive.org
When disturbed, individuals give a 'cough' alarm call and the herd retreats into thick bamboo thickets and lies on the ground for camouflage.Huffman, Brent

''Ultimate Ungulate'' via arkive.org
Takin feed in the early morning and late afternoon, grazing on a variety of leaves and grasses, as well as bamboo shoots and flowers. They have been observed standing on their hind legs to feed on leaves over high. Salt is also an important part of their diets, and groups may stay at a mineral deposit for several days.


Threats

The takin is listed as Vulnerable on the IUCN Red List and considered Endangered in China. It is threatened by overhunting and the destruction of its natural habitat. It is not a common species naturally, and the population appears to have been reduced considerably. Takin horns have appeared in the illegal wildlife trade in
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
; and during three surveys carried out from 1999 to 2006 in the Tachilek market, a total of 89 sets of horns were observed openly for sale.


Taxonomy

Relationships with other caprines based on mitochondrial DNA: In 2022, researchers found that the takin likely represents two species, the Himalayan takin (''B. taxicolor'') and the Chinese takin (''B. tibetana'').


References

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Further reading

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External links

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Wild creamy-white ''golden takins'' (b. t. bedfordi) in China
{{Taxonbar, from1=Q193469, from2=Q10741556 Caprids Mammals described in 1850 Taxa named by Brian Houghton Hodgson