Scabiosa
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''Scabiosa'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
in the honeysuckle family (
Caprifoliaceae The Caprifoliaceae or honeysuckle family is a clade of dicotyledonous flowering plants consisting of about 860 species, in 33, to 42 genera, with a nearly cosmopolitan distribution. Centres of diversity are found in eastern North America an ...
) of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
s. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in related genera such as ''
Knautia ''Knautia'' is a genus of flowering plants in the family Caprifoliaceae The Caprifoliaceae or honeysuckle family is a clade of dicotyledonous flowering plants consisting of about 860 species, in 33, to 42 genera, with a nearly cosmopolitan ...
'' and '' Succisa''; at least some of these were formerly placed in ''Scabiosa''. Another common name for members of this genus is pincushion flowers.


Etymology

The common name 'scabious' comes the Latin word ''scabiosus'' meaning 'mangy, rough or itchy' which refers to the herb's traditional usage as a
folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) comprises medical aspects of traditional knowledge that developed over generations within the folk beliefs of various societies, including indigenous peoples, before the ...
to treat scabies, an illness that causes a severe itching sensation.


Description

Some species of ''Scabiosa'' are
annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year ** Yearbook ** Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), ...
s, others
perennials A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
. Some are
herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
s; others have woody
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
s. The leaves of most species are somewhat hairy and partly divided into lobes, but a few are smooth and some species have simple leaves. The flowers are borne on
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed o ...
s in the form of heads; each head contains many small florets, each floret cupped in a membranous, saucer-shaped bract. The calyx has five
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 The term ''sepalum'' was coined b ...
s in the form of awns almost as long as the petals. After the flowers have dropped, the calyces together with the bracts form a spiky ball that may be the reason for the "pincushion" common name. The calyx is persistent and remains as a crown on the fruit after it is shed. The corolla has four to five lobes fringing a narrow funnel with a furry throat, the funnel being somewhat longer than the lobes. The florets have four stamens each, set high in the tube, and sticking out. Each fruit has just one seed. In a few species the heads are
sessile Sessility, or sessile, may refer to: * Sessility (motility), organisms which are not able to move about * Sessility (botany), flowers or leaves that grow directly from the stem or peduncle of a plant * Sessility (medicine), tumors and polyps that ...
but in most species they are borne singly on a tall peduncle. ''Scabiosa'' species and varieties differ in the colours of their flowers, but most are soft lavender blue, lilac or creamy white.


Taxonomy

It was first published in
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
on page 98 in 1753.


Species

As accepted by Plants of the World Online; * ''
Scabiosa adzharica ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa africana ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa albanensis ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa amoena ''Scabiosa columbaria'', called the small scabious or dwarf pincushion flower, is a widespread species of flowering plant in the genus ''Scabiosa'', native to Europe, Africa, and western Asia, from Sweden to Angola. In the garden it is a short-li ...
'' * ''
Scabiosa andryifolia ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa angustiloba ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa arenaria ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in ...
'' * ''
Scabiosa atropurpurea ''Scabiosa atropurpurea'' ( syn. ''Sixalix atropurpurea''), the mourningbride, mournful widow, pincushion flower, or sweet scabious, is an ornamental plant of the genus ''Scabiosa'' in the family Caprifoliaceae. It is native to southern Europe. ...
'' * '' Scabiosa austroafricana'' * '' Scabiosa balcanica'' * '' Scabiosa × beauverdiana'' * '' Scabiosa bipinnata'' * '' Scabiosa buekiana'' * '' Scabiosa canescens'' * '' Scabiosa cartenniana'' * '' Scabiosa cephalarioides'' * '' Scabiosa cinerea'' * '' Scabiosa colchica'' * ''
Scabiosa columbaria ''Scabiosa columbaria'', called the small scabious or dwarf pincushion flower, is a widespread species of flowering plant in the genus ''Scabiosa'', native to Europe, Africa, and western Asia, from Sweden to Angola. In the garden it is a short-li ...
'' * '' Scabiosa comosa'' * ''
Scabiosa correvoniana ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa corsica'' * ''
Scabiosa crinita ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa daucoides'' * '' Scabiosa drakensbergensis'' * '' Scabiosa eremophila'' * '' Scabiosa farinosa'' * '' Scabiosa fumarioides'' * ''
Scabiosa galianoi ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa holosericea'' * '' Scabiosa imeretica'' * '' Scabiosa incisa'' * '' Scabiosa ispartaca'' * '' Scabiosa japonica'' * '' Scabiosa jezoensis'' * '' Scabiosa lacerifolia'' * '' Scabiosa lachnophylla'' * '' Scabiosa libyca'' * ''
Scabiosa lucida ''Scabiosa lucida'', commonly known as shining scabious, is a species of flowering plants in the honeysuckle family (Caprifoliaceae). Taxonomy ''Scabiosa lucida'' contains the following subspecies: * ''Scabiosa lucida stricta'' * ''Scabiosa luci ...
'' * '' Scabiosa × lucidula'' * '' Scabiosa mollissima'' * '' Scabiosa nitens'' * ''
Scabiosa ochroleuca ''Scabiosa ochroleuca'', commonly called cream pincushions or cream scabious, is a species of scabious ''Scabiosa'' is a genus in the honeysuckle family ( Caprifoliaceae) of flowering plants. Many of the species in this genus have common n ...
'' * ''
Scabiosa owerinii ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa paphlagonica'' * '' Scabiosa parielii'' * '' Scabiosa parviflora'' * '' Scabiosa praemontana'' * '' Scabiosa pyrenaica'' * '' Scabiosa semipapposa'' * '' Scabiosa silenifolia'' * '' Scabiosa sirnakia'' * '' Scabiosa sivrihisarica'' * ''
Scabiosa solymica ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa sosnowskyi'' * '' Scabiosa taygetea'' * '' Scabiosa tenuis'' * '' Scabiosa thysdrusiana'' * '' Scabiosa transvaalensis'' * '' Scabiosa triandra'' * '' Scabiosa triniifolia'' * '' Scabiosa turolensis'' * ''
Scabiosa tuzluca ''Scabiosa'' is a genus in the honeysuckle family (Caprifoliaceae) of flowering plants. Many of the species in this genus have common names that include the word scabious, but some plants commonly known as scabious are currently classified in r ...
'' * '' Scabiosa tysonii'' * '' Scabiosa velenovskiana'' * '' Scabiosa vestina'' * '' Scabiosa webbiana''


Distribution

Members of this genus are native to Africa, Europe and Asia. Some species of ''Scabiosa'', notably
small scabious ''Scabiosa columbaria'', called the small scabious or dwarf pincushion flower, is a widespread species of flowering plant in the genus ''Scabiosa'', native to Europe, Africa, and western Asia, from Sweden to Angola. In the garden it is a short-li ...
(''S. columbaria'') and Mediterranean sweet scabious (''S. atropurpurea'') have been developed into cultivars for gardeners. In 1782, a mysterious pale yellow scabious, called '' Scabiosa trenta'', was described by
Belsazar Hacquet Belsazar de la Motte Hacquet (also Balthasar or Balthazar Hacquet) (c. 1739 – 10 January 1815) was a Carniolan physician of French descent in the Enlightenment Era. He was a war surgeon, a surgeon in the mining town of Idrija, and a professor o ...
, an Austrian physician, botanist, and mountaineer, in his work '' Plantae alpinae Carniolicae''. It became a great source of inspiration for later botanists and mountaineers discovering the
Julian Alps The Julian Alps ( sl, Julijske Alpe, it, Alpi Giulie, , ) are a mountain range of the Southern Limestone Alps that stretch from northeastern Italy to Slovenia, where they rise to 2,864 m at Mount Triglav, the highest peak in Slovenia. A large p ...
, especially
Julius Kugy Julius Kugy (19 July 1858 – 5 February 1944) was a mountaineer, writer, botanist, humanist, lawyer and officer of Slovenian descent. He wrote mostly in German. He is renowned for his travelogues from opening up the Julian Alps, in which he refle ...
. The Austrian botanist
Anton Kerner von Marilaun Anton Kerner Ritter von Marilaun, or Anton Joseph Kerner, (12 November 1831 – 21 June 1898) was an Austrian botanist and professor at the University of Vienna. Career Kerner was born in Mautern, Lower Austria, and studied medicine in Vienna f ...
later proved Belsazar Hacquet had not found a new species, but a specimen of the already known submediterranean ''
Cephalaria leucantha ''Cephalaria leucantha'' is a species of flowering plants in the family Caprifoliaceae. Description ''Cephalaria leucantha'' grows to in height. This hardy perennial plant has a long stem with divided, fern-like leaves. It produces white or pal ...
''.


Ecology

Scabious flowers are nectar rich and attractive to many
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...
s including
butterflies Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
and
moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s such as the
six-spot burnet The six-spot burnet (''Zygaena filipendulae'') is a day-flying moth of the family Zygaenidae. Subspecies *''Z. f. altapyrenaica'' Le Charles, 1950 *''Z. f. arctica'' Schneider, 1880 *''Z. f. balcanirosea'' Holik, 1943 *''Z. f. campaniae'' Re ...
. ''Scabiosa'' species are food plants for the
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e of some species of
Lepidoptera Lepidoptera ( ) is an order (biology), order of insects that includes butterfly, butterflies and moths (both are called lepidopterans). About 180,000 species of the Lepidoptera are described, in 126 Family (biology), families and 46 Taxonomic r ...
such as the
grey pug The grey pug (''Eupithecia subfuscata'') is a moth of the family Geometridae. It is found throughout the Palearctic region. It is also found in North America. Since it does not place any special demands on climatic conditions, special caterpilla ...
moth.


References

{{Taxonbar, from=Q147772 Caprifoliaceae genera Taxa named by Carl Linnaeus