Symplocos Costata
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''Symplocos'' is a genus of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s in the order
Ericales The Ericales are a large and diverse order of flowering plants in the asterid group of the eudicots. Well-known and economically important members of this order include tea and ornamental camellias, persimmon, ebony, blueberry, cranberry, l ...
. It contains about 300 species distributed in Asia and the Americas. Many species grow in humid tropical regions. This is sometimes considered to be the only genus in family
Symplocaceae Symplocaceae is a family of flowering plants in the order Ericales The Ericales are a large and diverse order of flowering plants in the asterid group of the eudicots. Well-known and economically important members of this order include tea ...
. Plants in this family are
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s and trees with white or yellow flowers. The oldest fossils of the genus date to the lower
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of Europe and North America, with the genus being present in Europe as late as the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
seeds of †''Symplocos granulosa'' are frequent in sediment rock layers of the
Late Oligocene The Chattian is, in the geologic timescale The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the pro ...
to the
Late Miocene The Late Miocene (also known as Upper Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages. The Tortonian and Messinian stages comprise the Late Miocene sub-epoch, which lasted from 11.63 Ma (million ye ...
of
Denmark Denmark is a Nordic countries, Nordic country in Northern Europe. It is the metropole and most populous constituent of the Kingdom of Denmark,, . also known as the Danish Realm, a constitutionally unitary state that includes the Autonomous a ...
,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
,
Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
and
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
. The fossil seeds are very similar to the seeds of the extan southern Chinese species ''Symplocos glandulifera'' and ''Symplocos sulcata''. Fossil seeds of †''Symplocos paucicostata'' are known from the Middle Pliocene sediment rock layers in Reuver, the
Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...
and from the
Late Pliocene Late or LATE may refer to: Everyday usage * Tardy, or late, not being on time * Late (or the late) may refer to a person who is dead Music * Late (The 77s album), ''Late'' (The 77s album), 2000 * Late (Alvin Batiste album), 1993 * Late!, a pseudo ...
sediment rock layers in
northern Italy Northern Italy (, , ) is a geographical and cultural region in the northern part of Italy. The Italian National Institute of Statistics defines the region as encompassing the four Northwest Italy, northwestern Regions of Italy, regions of Piedmo ...
. The fossil seeds are very similar to the seeds of the extant
East Asian East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
species '' Symplocos paniculata''.


Selected species

* '' Symplocos adenophylla'' * '' Symplocos ampulliformis'' — Northeast Queensland, Australia * '' Symplocos anamallayana'' * '' Symplocos anomala'' * '' Symplocos austromexicana'' — deciduous shrub up to 2m; narrow
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
,
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
* '' Symplocos badia'' * '' Symplocos baehnii'' * '' Symplocos barberi'' * '' Symplocos bauerlenii'' — shrub or small tree up to 7m; eastern
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
* '' Symplocos blancae'' * '' Symplocos bractealis'' * '' Symplocos breedlovei'' * '' Symplocos calycodactylos'' * '' Symplocos candelabrum'' — tree up to 13m;
Lord Howe Island Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland Port ...
* '' Symplocos canescens'' * '' Symplocos carmencitae'' * '' Symplocos chloroleuca'' * '' Symplocos clethrifolia'' * '' Symplocos citrea'' * '' Symplocos coccinea'' * ''
Symplocos cochinchinensis ''Symplocos cochinchinensis'' is a species of flowering plant in the sapphire-berry family Symplocaceae, native to tropical and subtropical Asia. The widespread ''Symplocos cochinchinensis'' subsp./var. ''laurina'' is now considered a synonym of ...
'' :* var. ''cochinchinensis'' :* var. ''gittonsii'' — Northeast Queensland, Australia :* var. ''glaberrima'' — Northeast Queensland, Australia :* var. ''pilosiuscula'' — Northeast Queensland, Australia * '' Symplocos cordifolia'' —
Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
* '' Symplocos coreana'' — Korean sweetleaf, deciduous shrub up to 5m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
* '' Symplocos coronata'' — Sri Lanka * '' Symplocos costaricana'' – Central America * '' Symplocos costata'' * '' Symplocos crassiramifera'' — Northeast Queensland, Australia * '' Symplocos cuneata'' — Sri Lanka * '' Symplocos cyanocarpa'' — Northeast Queensland, Australia * '' Symplocos ecuadorica'' – Ecuador * ''Symplocos foliosa'' - Endemic to the Western Ghats of India * '' Symplocos fuscata'' * '' Symplocos glauca'' — evergreen tree up to 15m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Indochina Mainland Southeast Asia (historically known as Indochina and the Indochinese Peninsula) is the continental portion of Southeast Asia. It lies east of the Indian subcontinent and south of Mainland China and is bordered by the Indian Ocean to th ...
* '' Symplocos graniticola'' — Northeast Queensland, Australia * '' Symplocos hayesii'' — Northeast Queensland, Australia * '' Symplocos henschelii'' * '' Symplocos hispidula'' * '' Symplocos hylandii'' — Northeast Queensland, Australia * '' Symplocos junghuhnii'' * †'' Symplocos kowalewskii''
Baltic amber Baltic amber or succinite is amber from the Baltic region, home of its largest known deposits. It was produced sometime during the Eocene epoch, but exactly when is controversial. It has been estimated that this forested region provided the re ...
,
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
* '' Symplocos lancifolia'' — evergreen tree up to 5m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
* '' Symplocos longipes'' * ''
Symplocos lucida ''Symplocos'' is a genus of flowering plants in the order Ericales. It contains about 300 species distributed in Asia and the Americas. Many species grow in humid tropical regions. This is sometimes considered to be the only genus in family Sympl ...
'' —
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
, South
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
,
Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms. It is a phytogeographical floristic region in the Paleotropical kingdom. It was first recognized as a distinct region ...
* '' Symplocos lugubris'' * '' Symplocos mezii'' * '' Symplocos myrtacea'' — evergreen tree up to 10m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
* '' Symplocos nairii'' * '' Symplocos nicolsonii'' —
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
* '' Symplocos nivea'' * '' Symplocos octopetala'' * '' Symplocos oligandra'' * '' Symplocos paniculata'' — (sapphireberry) — deciduous shrub up to 8m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
; popular as an ornamental plant * '' Symplocos paucistaminea'' — Northeast Queensland, Australia * '' Symplocos pendula'' * '' Symplocos peruviana'' * '' Symplocos pluribracteata'' * '' Symplocos prunifolia'' — evergreen tree]up to 10m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
* '' Symplocos pulchra'' — Sri Lanka * '' Symplocos pyriflora'' * '' Symplocos racemosa'' —
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
, South
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
* '' Symplocos rimbachii'' * '' Symplocos shilanensis'' * '' Symplocos sousae'' * '' Symplocos stawellii'' — shrub or small tree up to 30m; eastern
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
:* var. ''montana'' — Northeast Queensland, Australia :* var. ''stawellii'' — Australia * '' Symplocos subandina'' * '' Symplocos subintegra''
Chatterjee Chatterjee (), also known as Chattopadhyay is a Bengali Hindu surname, used by the Kulin group of the Bengali Brahmin caste. Together with Banerjees, Mukherjees, Bhattacharjees, and Gangulys, Chatterjees form the Kulin Brahmins, the highest ...
* '' Symplocos tacanensis'' * '' Symplocos tanakae'' — evergreen tree;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
* '' Symplocos theophrastaefolia'' — evergreen tree up to 15m;
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
* '' Symplocos thwaitesii'' — shrub or small tree up to 17m; eastern
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
* '' Symplocos tinctoria'' — (sweetleaf, horse-sugar, yellowwood) — deciduous or evergreen shrub or tree up to 6m;
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
* '' Symplocos trichoclada'' * '' Symplocos truncata'' * '' Symplocos tubulifera'' * '' Symplocos verrucisurcula'' * '' Symplocos versicolor''


References

Ericales genera {{Ericales-stub