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''Pinus cembra'', also known as Swiss pine, Swiss stone pine or Arolla pine or Austrian stone pine or just stone pine, is a species of a pine tree in the subgenus ''Strobus''.


Description

The Swiss pine is a member of the white pine group, ''Pinus'' subgenus ''Strobus'', and like all members of that group, the
leaves A leaf (plural, : leaves) is any of the principal appendages of a vascular plant plant stem, stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", wh ...
('needles') are in fascicles (bundles) of five, with a deciduous sheath. The mature size is typically between and in height, and the trunk diameter can be up to . However, it grows very slowly and it may take 30 years for the tree to reach . The cones, which contain the seeds (or nuts), of the Swiss pine are to long. Cones take 2 years (24 months) to mature. The to long seeds have only a vestigial wing and are dispersed by spotted nutcrackers. The species is long-lasting and can reach an age between 500 and 1000 years. In its natural environment, this tree usually reaches
reproductive maturity Sexual maturity is the capability of an organism to reproduce. In humans it might be considered synonymous with adulthood, but here puberty is the name for the process of biological sexual maturation, while adulthood is based on cultural definiti ...
at the age of 50 years (if the conditions are more extreme, even 80 years). The very similar Siberian pine (''Pinus sibirica'') is treated as a variety or subspecies of Swiss pine by some botanists. The Siberian pine differs in having slightly larger cones, being more massive (taller, wider trunk), having a faster growth rate and needles with three resin canals instead of two as in the Swiss pine. File:Pinus cembra cones in Gröden crop.jpg, Cones of ''Pinus cembra'' File:Fresh Pinus cembra cone in June.jpg, Close-up of an immature cone from a cultivated ''Pinus cembra'' File:Pinus cembra cone dried.jpg, Close-up of a mature cone from a cultivated ''Pinus cembra'' File:Pinus cembra seedling planted for pine nut production.jpg, 5-year-old ''Pinus cembra'' seedling planted for pine nut production, Baldone,
Latvia Latvia ( or ; lv, Latvija ; ltg, Latveja; liv, Leţmō), officially the Republic of Latvia ( lv, Latvijas Republika, links=no, ltg, Latvejas Republika, links=no, liv, Leţmō Vabāmō, links=no), is a country in the Baltic region of ...
File:Bark of an young Arolla pine.jpg,
Bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, ...
of a young ''Pinus cembra'',
National Botanic Garden of Latvia The National Botanic Garden of Latvia ( lv, Nacionālais botāniskais dārzs) is a botanical garden in Salaspils, Latvia. It is one of the largest botanical gardens in the Baltic states. History The present institution was founded in 1956, but ...
File:Pinus cembra coupe MHNT.jpg, Cross-section of ''Pinus cembra'' - 181 years - MHNT File:Swiss pine (Pinus cembra) 'Columnaris' cone.jpg, ''Pinus cembra'' 'Columnaris' (cultivar) 1 year-old cone and foliage


Distribution and habitat

The Swiss pine grows in the Alps and
Carpathian Mountains The Carpathian Mountains or Carpathians () are a range of mountains forming an arc across Central Europe. Roughly long, it is the third-longest European mountain range after the Urals at and the Scandinavian Mountains at . The range stretches ...
of central Europe, in Poland ( Tatra Mountains),
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
, France, Italy, Austria, Germany, Slovenia, Slovakia ( Tatra Mountains), Ukraine and Romania. It typically grows at to altitude. It often reaches the alpine tree line in this area.


Ecology

Swiss pine associates with numerous species of mycorrhizal fungi from a young age, usually from the genus '' Suillus''. This
symbiosis Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit ...
improves the tree's growth and survival rate.


Uses

Swiss pine is a popular ornamental tree in
park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are urban green space, green spaces set aside for recreation inside t ...
s and large gardens, giving steady though not fast growth on a wide range of sites where the climate is cold. It is very tolerant of severe winter cold, hardy down to at least , and also of wind exposure. The seeds are also harvested and sold as pine nuts. When cultivated, it will likely start producing cones after the age of 12 years, much faster than in the wild. This depends on the climate, soil type,
mycorrhiza   A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant ...
l fungi etc. To make the tree bear cones faster, the tree can be inoculated with
ectomycorrhiza An ectomycorrhiza (from Greek ἐκτός ', "outside", μύκης ', "fungus", and ῥίζα ', "root"; pl. ectomycorrhizas or ectomycorrhizae, abbreviated EcM) is a form of symbiotic relationship that occurs between a fungal symbiont, or mycobi ...
l fungi, such as '' Suillus luteus, Suillus americanus, Suillus placidus'' etc. These are the most effective. Pine cones cut into slices are used to flavor schnapps, which is then sold as "Zirbenschnaps" or "Zirbeler" schnapps. The wood is the most used for carvings in Val Gardena since the 17th century. The cone of the Swiss pine was the field sign of the Roman legion stationed in
Rhaetia Raetia ( ; ; also spelled Rhaetia) was a province of the Roman Empire, named after the Rhaetian people. It bordered on the west with the country of the Helvetii, on the east with Noricum, on the north with Vindelicia, on the south-west with Tr ...
in 15 BC, and hence it is used as the heraldic charge (known as ''Zirbelnuss'' in German) in the coat of arms of the city of Augsburg, the site of the Roman fort Augusta Vindelicorum. It is also a species that is often used in bonsai. ''Pinus cembra'' can be found in the uppermost forest belt where it helps to minimize the risk of avalanches and soil erosion. Due to this ability, the tree is valued as a stabilizing factor for afforestation projects at high elevations. Like other European and Asian white pines, Swiss pine is very resistant to white pine blister rust (''Cronartium ribicola''). This fungal disease was accidentally introduced from Europe into
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car ...
, where it has caused severe mortality in the American native white pines in many areas, notably, Western white pine and the closely related whitebark pine. Swiss pine is of great value for research into hybridization to develop rust resistance in these species.


References


External links


''Pinus cembra'' cone pic (scroll to bottom of page)



Link to Joanneum Research Study

Folder Joanneum Research: Stone Pine - Positive health effects of Stone Pine furniture

''Pinus cembra''
- distribution map, genetic conservation units and related resources. European Forest Genetic Resources Programme (EUFORGEN) {{Authority control Edible nuts and seeds Flora of Poland Least concern plants Cembra Plants described in 1753 Taxa named by Carl Linnaeus Trees of Europe Flora of France Flora of the Alps Flora of the Carpathians