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''Strychnos'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s, belonging to the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Loganiaceae The Loganiaceae are a family of flowering plants classified in order Gentianales. The family includes up to 13 genera, distributed around the world's tropics. There are not any great morphological characteristics to distinguish these taxa from ot ...
(sometimes Strychnaceae). The genus includes about 200 accepted
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s and
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
s. The genus is widely distributed around the world's tropics and is noted for the presence of poisonous indole alkaloids in the roots, stems and leaves of various species. Among these alkaloids are the well-known and virulent poisons
strychnine Strychnine (, , American English, US chiefly ) is a highly toxicity, toxic, colorless, bitter, crystalline alkaloid used as a pesticide, particularly for killing small vertebrates such as birds and rodents. Strychnine, when inhaled, swallowed, ...
and
curare Curare ( or ; or ) is a common name for various alkaloid arrow poisons originating from plant extracts. Used as a paralyzing agent by indigenous peoples in Central and South America for hunting and for therapeutic purposes, curare only ...
.


Etymology

The name ''strychnos'' was applied by
Pliny the Elder Gaius Plinius Secundus (AD 23/24 79), known in English as Pliny the Elder ( ), was a Roman Empire, Roman author, Natural history, naturalist, and naval and army commander of the early Roman Empire, and a friend of the Roman emperor, emperor Vesp ...
in his ''
Natural History Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
'' to ''
Solanum nigrum ''Solanum nigrum'', the European black nightshade or simply black nightshade or blackberry nightshade, is a species of flowering plant in the family Solanaceae, native to Eurasia and introduced in the Americas, Australasia, and South Africa. Ripe ...
''. The word is derived from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
στρύχνον (''strúkhnon'') – "acrid", "bitter". The meaning of the word ''strychnos'' was not fixed in Ancient Greece, where it could designate a variety of different plants having in common the property of toxicity.


Distribution

The genus has a
pantropical A pantropical ("all tropics") distribution is one which covers tropical regions of both the Eastern and Western hemispheres. Examples of species include caecilians, modern sirenians and the plant genera ''Acacia'' and ''Bacopa''. ''Neotropical' ...
distribution.


Taxonomy

The genus is divided into 12 sections, though it is conceded that the sections do not reflect evolution of the genus, and all sections except ''Spinosae'' are polyphyletic: * ''Strychnos'' (53 species) * ''Rouhamon'' (21 species) * ''Breviflorae'' (32 species) * ''Penicillatae'' (17 species) * ''Aculeatae'' (1 species) * ''Spinosae'' (4 species) * ''Brevitubae'' (18 species) * ''Lanigerae'' (32 species) * ''Phaeotrichae'' (1 species) * ''Densiflorae'' (8 species) * ''Dolichantae'' (9 species) * ''Schyphostrychnos'' (1 species)


Selected species

*''
Strychnos benthami ''Strychnos benthami'' is a species of plant in the Loganiaceae family. It is endemic to Sri Lanka Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia ...
'' C.B.Clarke *''
Strychnos camptoneura ''Strychnos camptoneura'' is a species of plant in the Loganiaceae family. It is native to Cameroon, the Central African Republic, the Democratic Republic of the Congo, Gabon, Ivory Coast, Liberia, Nigeria, the Republic of the Congo and Zaire. D ...
'' Gilg &
Busse Busse is a surname. People with this surname include: *Andreas Busse (born 1959), East German (now German) former middle-distance runner *Carl Busse (architect) (1834-1896), German architect and master builder *Carl Hermann Busse (1872–1918), Ger ...
*''
Strychnos chromatoxylon ''Strychnos chromatoxylon'' is a species of plant in the Loganiaceae family. It is found in Cameroon, Central African Republic, and Ivory Coast Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a ...
'' Leeuwenb. *''
Strychnos cocculoides ''Strychnos cocculoides'', also known as the corky-bark monkey orange tree, or ''Ntonga'', and ''suurklapper'' in Afrikaans, is a fruiting tree of Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed ...
''
Baker A baker is a tradesperson who baking, bakes and sometimes Sales, sells breads and other products made of flour by using an oven or other concentrated heat source. The place where a baker works is called a bakery. History Ancient histo ...
*''
Strychnos elaeocarpa ''Strychnos elaeocarpa'' is a species of plant in the Loganiaceae family. It is endemic to Cameroon. Its natural habitat is subtropical or tropical moist lowland forests. It is threatened by habitat loss Habitat destruction (also termed h ...
'' Gilg ex Leeuwenb. *''
Strychnos icaja ''Strychnos icaja'' is a species belonging to the plant family Loganiaceae, native to West Tropical Africa. It is a very large, tropical rainforest liana which may attain a length of . Taxonomy The species was published in the journal ''Adans ...
'' Baill. *''
Strychnos ignatii ''Strychnos ignatii'' is a tree in the family Loganiaceae, native to the Philippines, particularly in Catbalogan and parts of China. The plant was first described by the Moravian (Czech) Jesuit working in the Philippines, brother Georg Kamel ...
'' P.J. Bergius *'' Strychnos madagascariensis'' Poir. *'' Strychnos mellodora'' S.Moore *'' Strychnos millepunctata'' Leeuwenb. *'' Strychnos nux-blanda'' A.W.Hill *''
Strychnos nux-vomica ''Strychnos nux-vomica'', the strychnine tree, also known as nux vomica, poison fruit, semen strychnos, and quaker buttons, is a deciduous tree native to India and to southeast Asia. It is a medium-sized tree in the family Loganiaceae that grows ...
'' L. *'' Strychnos potatorum'' L.f. *'' Strychnos psilosperma''
F.Muell. Baron Sir Ferdinand Jacob Heinrich von Mueller, (; 30 June 1825 – 10 October 1896) was a German-Australian physician, geographer, and most notably, a botanist. He was appointed government botanist for the then colony of Victoria (state), Vic ...
*'' Strychnos pungens'' Soler. *'' Strychnos spinosa'' Lam. *'' Strychnos staudtii'' Gilg *'' Strychnos tetragona'' A.W.Hill *''
Strychnos toxifera ''Strychnos toxifera'', called bush rope and devil doer, is a species of flowering plant in the genus ''Strychnos'', native to Costa Rica, Panama, Colombia, Ecuador, Venezuela, the Guianas, Brazil, Peru and Bolivia. It is the principal source of ...
'' R.H.Schomb. ex Lindl. *'' Strychnos usambarensis'' Gilg ex
Engl. Engl or Engl. may refer to: *England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more ...
* The
strychnine tree ''Strychnos nux-vomica'', the strychnine tree, also known as nux vomica, poison fruit, semen strychnos, and quaker buttons, is a deciduous tree native to India and to southeast Asia. It is a medium-sized tree in the family Loganiaceae that grows ...
, ''Strychnos nux-vomica'', native to tropical Asia, is the source of the poison
strychnine Strychnine (, , American English, US chiefly ) is a highly toxicity, toxic, colorless, bitter, crystalline alkaloid used as a pesticide, particularly for killing small vertebrates such as birds and rodents. Strychnine, when inhaled, swallowed, ...
. * ''Strychnos tonga'', native to
Tonga Tonga, officially the Kingdom of Tonga, is an island country in Polynesia, part of Oceania. The country has 171 islands, of which 45 are inhabited. Its total surface area is about , scattered over in the southern Pacific Ocean. accordin ...
, is a synonym of ''Strychnos spinosa''. * ''
Strychnos ignatii ''Strychnos ignatii'' is a tree in the family Loganiaceae, native to the Philippines, particularly in Catbalogan and parts of China. The plant was first described by the Moravian (Czech) Jesuit working in the Philippines, brother Georg Kamel ...
'' ("St. Ignatius bean"), is a closely related Asian shrub/tree. * The species ''
Strychnos toxifera ''Strychnos toxifera'', called bush rope and devil doer, is a species of flowering plant in the genus ''Strychnos'', native to Costa Rica, Panama, Colombia, Ecuador, Venezuela, the Guianas, Brazil, Peru and Bolivia. It is the principal source of ...
'' is a principal plant source of the
arrow poison Arrow poisons are used to poison arrow heads or darts for the purposes of hunting and warfare. They have been used by indigenous peoples worldwide and are still in use in areas of South America, Africa and Asia. Notable examples are the poisons se ...
curare Curare ( or ; or ) is a common name for various alkaloid arrow poisons originating from plant extracts. Used as a paralyzing agent by indigenous peoples in Central and South America for hunting and for therapeutic purposes, curare only ...
. * Three trees from
Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
, commonly known as " monkey oranges", are drought-tolerant and produce popular edible fruits: the corky-barked monkey orange or suurklapper, ''
Strychnos cocculoides ''Strychnos cocculoides'', also known as the corky-bark monkey orange tree, or ''Ntonga'', and ''suurklapper'' in Afrikaans, is a fruiting tree of Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed ...
''; the Natal orange or green or spiny monkey orange, '' Strychnos spinosa''; and the black or spiny-leaved monkey orange '' Strychnos pungens''. * The ripe seeds of '' Strychnos potatorum'', known as Therran or Nirmal, can be ground and used as a coagulant to purify water; or they may be rubbed against the inside walls of the earthenware water containers. Mrs Grieve's Herbal of 1931 also mentions traditional water purification uses of an Indian species called ''Strychnos pseudo'' (not a valid botanical name). *Two very well preserved fossilised corollas with
stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s and styles from flowers of a plant that has been named ''Strychnos electri'' (the Latin name of amber is ''electrum''), believed to be a vine, were discovered in
amber Amber is fossilized tree resin. Examples of it have been appreciated for its color and natural beauty since the Neolithic times, and worked as a gemstone since antiquity."Amber" (2004). In Maxine N. Lurie and Marc Mappen (eds.) ''Encyclopedia ...
from the Dominican Republic. The amber is from tropical tree '' Hymenaea protera'', formerly abundant but now extinct, which formed part of the forest canopy. The age of the amber is believed to be between 15 and 45 million years, from the mid-
Tertiary period The Tertiary ( ) is an obsolete Period (geology), geologic period spanning 66 million to 2.6 or 1.8 million years ago. The period began with the extinction of the non-bird, avian dinosaurs in the Cretaceous–Paleogene extinction event, at t ...
. This demonstrates an early date for these plants.New species of ancient tropical flower found in amber from the Dominican Republic
Rachel Sullivan,
ABC News Online ABC News, also known as ABC News and Current Affairs, is a public news service produced by the Australian Broadcasting Corporation. The service covers both local and world affairs, broadcasting both nationally as ABC News, and across the Asia- ...
, 16 February 2016
, Article number 16005


Gallery

File:Strychnos madagascariensis (4337703118).jpg, '' Strychnos madagascariensis'' File:Strychnos nux-vomica flowers 04.JPG, ''
Strychnos nux-vomica ''Strychnos nux-vomica'', the strychnine tree, also known as nux vomica, poison fruit, semen strychnos, and quaker buttons, is a deciduous tree native to India and to southeast Asia. It is a medium-sized tree in the family Loganiaceae that grows ...
'' flowers File:Strychnos potatorum 05.jpg, '' Strychnos potatorum'' File:Strychnos psilosperma foliage and fruit.jpg, '' Strychnos psilosperma'' File:Strychnos pungens, blomme, b, Seringveld.jpg, '' Strychnos pungens'' flowers File:Strychnos pungens, vrug, Little Eden.jpg, '' Strychnos pungens'' detached fruit File:Strychnos spinosa tree.jpg, '' Strychnos spinosa'' tree in fruit File:Strychnos usambarensis00.jpg, '' Strychnos usambarensis'' in flower File:Strychnos usambarensis02.jpg, '' Strychnos usambarensis'' in fruit


See also

* List of Southern African indigenous trees and woody lianes


References


External links

* {{Authority control Gentianales genera Pantropical flora