A spiral valve or scroll valve is the corkscrew-shaped lower portion of the

intestine The gastrointestinal tract (GI tract, digestive tract, alimentary canal) is the tract or passageway of the digestive system that leads from the mouth to the anus. The tract is the largest of the body's systems, after the cardiovascular system. T ...

of some

shark Sharks are a group of elasmobranch cartilaginous fish characterized by a ribless endoskeleton, dermal denticles, five to seven gill slits on each side, and pectoral fins that are not fused to the head. Modern sharks are classified within the ...

Acipenseriformes Acipenseriformes is an order (biology), order of basal (phylogenetics), basal Actinopterygii, ray-finned fishes that includes living and fossil sturgeons and paddlefishes (Acipenseroidei), as well as the extinct family (biology), families C ...

(

sturgeon Sturgeon (from Old English ultimately from Proto-Indo-European language, Proto-Indo-European *''str̥(Hx)yón''-) is the common name for the 27 species of fish belonging to the family Acipenseridae. The earliest sturgeon fossils date to the ...

and

paddlefish Paddlefish (family Polyodontidae) are a family of ray-finned fish belonging to order Acipenseriformes, and one of two living groups of the order alongside sturgeons (Acipenseridae). They are distinguished from other fish by their elongated rost ...

), rays, skates,

bichir Bichirs and the reedfish comprise Polypteridae , a family (biology), family of archaic Actinopterygii, ray-finned fishes and the only family in the order (biology), order Polypteriformes .Helfman GS, Collette BB, Facey DE, Bowen BW. 2009. The D ...

s, Lepisosteiformes (gars), and

lungfish Lungfish are freshwater vertebrates belonging to the class Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, inc ...

es. A modification of the

ileum The ileum () is the final section of the small intestine in most higher vertebrates, including mammals, reptiles, and birds. In fish, the divisions of the small intestine are not as clear and the terms posterior intestine or distal intestine may ...

, the spiral valve is internally twisted or coiled to increase the surface area of the intestine which increases nutrient absorption.

Description

The intestines of a shark are much shorter than those of mammals. Sharks have compensated for this problem by having a spiral valve, or a scroll valve, inside the intestine to increase the absorbent surface of the intestine. By keeping digestible material in the ileum for an extended period maximum nutrient absorption is ensured. For this reason, many sharks and related

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

feed very infrequently. The food passes into the comparatively short colon of the shark almost fully digested, and then out the

cloaca A cloaca ( ), : cloacae ( or ), or vent, is the rear orifice that serves as the only opening for the digestive (rectum), reproductive, and urinary tracts (if present) of many vertebrate animals. All amphibians, reptiles, birds, cartilagin ...

and vent. A consequence of the spiral valve constricting the lumen of the ileum is that sharks cannot pass large hard objects (such as

bone A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...

s) through their lower intestine. Such objects instead remain in the

stomach The stomach is a muscular, hollow organ in the upper gastrointestinal tract of Human, humans and many other animals, including several invertebrates. The Ancient Greek name for the stomach is ''gaster'' which is used as ''gastric'' in medical t ...

until sufficiently broken down for passing through the valve region, or are regurgitated. Consequently, shark stomachs often contain items of interest that enable one to determine what the animals feed on, as well as non-food items ingested during a feeding frenzy. Sharks also possess a rectal gland that removes excess salt.

References

Fish anatomy Sharks

valve A valve is a device or natural object that regulates, directs or controls the flow of a fluid (gases, liquids, fluidized solids, or Slurry, slurries) by opening, closing, or partially obstructing various passageways. Valves are technically Pip ...

{{Shark-stub

Description

See also

References