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''Euonymus'' is a genus of flowering plants in the staff vine family
Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small trees, belonging to the order Celastrales. The great majority of the genera are tropical, with only ''Celastrus'' (the staf ...
. Common names vary widely among different species and between different English-speaking countries, but include spindle (or spindle tree), burning-bush, strawberry-bush, wahoo, wintercreeper, or simply euonymus. It has about 140 species''Euonymus''.
Flora of China.
Du, C., et al. (2013)
Revision of three species of ''Euonymus'' (Celastraceae) from China.
''Phytotaxa'' 109(1) 45-53.
of
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
and
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
shrubs, small trees and
liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...
s. They are mostly
native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...
to East Asia, extending to the Himalayas,''Botanica: The Illustrated A-Z of over 10000 Garden Plants and How to Cultivate Them''. Könemann, 2004. pg. 358. and they are also distributed in Europe, Australasia, North America, and Madagascar. Fifty species are
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to China.


Description

The inconspicuous flowers occur in small groups, and can be green, yellow, pink or maroon in color depending on species. The leaves are opposite (rarely alternate) and simple ovoid, typically 2–15 cm long, and usually with a finely serrated margin. The fruit is a pink or white four- or five-valved pod-like berry, which splits open to reveal the fleshy-coated orange or red
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s. The seeds are eaten by
frugivorous A frugivore ( ) is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance ...
birds, which digest the fleshy seed coat and disperse the seeds in their droppings. Many species are used in traditional medicines, and parts of the plants can be
poisonous A poison is any chemical substance that is harmful or lethal to living organisms. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figurati ...
to humans.


Cultivation and uses

The wood of some species was traditionally used to make spindles for
spinning Spin or spinning most often refers to: * Spin (physics) or particle spin, a fundamental property of elementary particles * Spin quantum number, a number which defines the value of a particle's spin * Spinning (textiles), the creation of yarn or thr ...
wool Wool is the textile fiber obtained from sheep and other mammals, especially goats, rabbits, and camelids. The term may also refer to inorganic materials, such as mineral wool and glass wool, that have some properties similar to animal w ...
; this use is the origin of the British English name of the shrubs. Euonymus are popular garden shrubs, grown for their foliage, the deciduous species often exhibiting very bright red autumnal colours, and also for the decorative berries. ''
Euonymus alatus ''Euonymus alatus'', known variously as burning bush, winged euonymus, winged spindle, and winged spindle-tree, is a species of flowering plant in the family Celastraceae, native to central and northern China, Japan, and Korea. It is a popular ...
'' (winged euonymus or burning-bush) is considered an invasive species in the woodlands of the northeastern United States.


Selected species

*'' Euonymus acanthocarpus'' *'' Euonymus acuminifolius'' *''
Euonymus alatus ''Euonymus alatus'', known variously as burning bush, winged euonymus, winged spindle, and winged spindle-tree, is a species of flowering plant in the family Celastraceae, native to central and northern China, Japan, and Korea. It is a popular ...
'' – winged spindle, burning-bush *''
Euonymus americanus ''Euonymus americanus'' is a species of flowering plant in the family Celastraceae. Common names include strawberry bush, American strawberry bush, bursting-heart, hearts-a-bustin, and hearts-bustin'-with-love. It is native to the eastern United ...
'' – strawberry-bush *''
Euonymus angulatus ''Euonymus angulatus'' is a species of tree in the family Celastraceae. They can grow up to tall and have dull purple flowers. They grow in medium elevation evergreen forests between . It is endemic to India, where it is known from Karnataka, K ...
'' *''
Euonymus atropurpureus ''Euonymus atropurpureus'' is a species of shrub in the bittersweet family. It has the common names American wahoo, eastern wahoo, burningbush and hearts bursting with love. It is native to eastern North America. Distribution and habitat This ...
'' – eastern burning-bush, eastern wahoo *''
Euonymus castaneifolius ''Euonymus castaneifolius'' is a tree in the family Celastraceae. The specific epithet ' is from the Latin meaning 'chestnut-coloured leaves'. Description ''Euonymus castaneifolius'' grows up to tall with a trunk diameter of up to . The smooth ...
'' *''
Euonymus carnosus ''Euonymus carnosus'', called the fleshyflowered spindletree, is a species of flowering plant in the genus ''Euonymus'', native to southeast and southcentral China, Taiwan, the Bonin Islands, the Ryukyu Islands, and Japan. It has gained the Royal ...
'' *''
Euonymus cochinchinensis ''Euonymus cochinchinensis'' is a tree of tropical Asia in the staff vine family Celastraceae. The specific epithet ' refers to the species being native to Indo-China. Description ''Euonymus cochinchinensis'' grows as a small tree up to tall. T ...
'' *''
Euonymus cornutus ''Euonymus cornutus'' is a species of flowering plant in the genus ''Euonymus'', native to Tibet, central China and Myanmar. Its putative variety ''Euonymus cornutus'' var. ''quinquecornutus'', called the fivehorned spindle, has gained the Royal ...
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Euonymus dichotomus ''Euonymus dichotomus'' is a species of flowering plant in the family Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small trees, belonging to the order Celast ...
'' *''
Euonymus dielsianus ''Euonymus dielsianus'' is a species of flowering plant in the family Celastraceae. It was first described by the German botanists Ludwig E. T. Loesener and Ludwig Diels Friedrich Ludwig Emil Diels (24 September 1874 – 30 November 1945) was ...
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Euonymus europaeus ''Euonymus europaeus'', the spindle, European spindle, or common spindle, is a species of flowering plant in the family Celastraceae, native to much of Europe, where it inhabits the edges of forest, hedges and gentle slopes, tending to thrive on ...
'' – European spindle *''
Euonymus fimbriatus ''Euonymus fimbriatus'', also known as the fringed spindle tree, is a plant from the genus ''Euonymus''. This tree comes from the Himalayas. This tree can grow up to tall and it is deciduous In the fields of horticulture and botany, the te ...
'' – fringed spindle tree *''
Euonymus fortunei ''Euonymus fortunei'', the spindle, Fortune's spindle, winter creeper or wintercreeper, is a species of flowering plant in the family Celastraceae, native to east Asia, including China, Korea, the Philippines and Japan. ''E. fortunei'' is highly ...
'' – Fortune's spindle, wintercreeper *''
Euonymus frigidus ''Euonymus frigidus'' (synonym ''Euonymus assamicus'') is a species of plant in the family Celastraceae. It is native to China, the Himalayas, Assam and Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referre ...
'' *''
Euonymus glandulosus ''Euonymus glandulosus'' is a species of plant in the family Celastraceae. It is a tree found in Borneo and the Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Sou ...
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Euonymus grandiflorus ''Euonymus grandiflorus'' is a species of plant in the family Celastraceae. It is found in China, Myanmar, and Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated ...
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Euonymus hamiltonianus ''Euonymus hamiltonianus'', known by the common names Hamilton's spindletree, Himalayan spindle, and Siebold's spindle is a species of flowering plant in the family Celastraceae. It is native to Asia, where it is distributed in Afghanistan, Russi ...
'' – Hamilton's spindle, Himalayan spindle, Siebold's spindle *''
Euonymus indicus ''Euonymus indicus'', the Indian spindle tree, is a small evergreen understorey tree in the family Celastraceae. It can grow up to a height of 13 m and girth up to 1 m. Description The leaves are simple and show opposite phyllotaxy. The petiol ...
'' *'' Euonymus japonicus'' – Japanese spindle, evergreen spindle *''
Euonymus lanceifolia ''Euonymus lanceifolia'' is a species of plant in the family Celastraceae. It is endemic to China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 bi ...
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Euonymus laxiflorus ''Euonymus laxiflorus'' (synonym ''Euonymus paniculatus'') is a species of plant in the family Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small trees, belo ...
'' *''
Euonymus lucidus ''Euonymus lucidus'' is a species of ''Euonymus'' native to the Himalaya region, from Pakistan east to northern Assam and Myanmar.Flora of Pakistan''Euonymus pendulus''/ref>Nepal Checklist''Euonymus pendulus''/ref> It is an evergreen shrub or sm ...
'' *'' Euonymus macropterus'' *'' Euonymus morrisonensis'' *''
Euonymus myrianthus ''Euonymus myrianthus'', the many-flowered spindle, is a species of flowering plant in the family Celastraceae. It is native to central and southern China. A variable evergreen shrub or small tree, and reaching in height, it is typically found i ...
'' – many-flowered spindle *'' Euonymus nanus'' – dwarf spindle *''
Euonymus obovatus ''Euonymus obovatus'', the running strawberry bush, is a trailing, woodland groundcover plant of the family Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small ...
'' – running strawberry-bush *'' Euonymus occidentalis'' – western burning-bush *'' Euonymus oxyphyllus'' – Korean spindletree *'' Euonymus phellomanus'' *'' Euonymus sachalinensis'' *''
Euonymus sanguineus ''Euonymus sanguineus'', the blood red spindle, is a species of flowering plant in the family Celastraceae The Celastraceae (staff-vine or bittersweet) are a family of 98 genera and 1,350 species of herbs, vines, shrubs and small trees, belong ...
'' – blood red spindle *'' Euonymus serratifolius'' *'' Euonymus verrucosus'' *'' Euonymus walkeri'' *'' Euonymus wilsonii'' – Chinese euonymus


References

{{Authority control Celastrales genera Plants described in 1753 Taxa named by Carl Linnaeus