The slender sunfish (''Ranzania laevis'') is a mola of the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Molidae The Molidae comprise the family of the molas or ocean sunfishes, unusual fish whose bodies come to an end just behind the dorsal fin, dorsal and anal fins, giving them a "half-fish" appearance. They are also the largest of the ray-finned bony fi ...

, the only extant member of the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

''Ranzania'',Matsuura, K. (2014): Taxonomy and systematics of tetraodontiform fishes: a review focusing primarily on progress in the period from 1980 to 2014. ''Ichthyological Research, 62 (1): 72–113.'' found globally in

tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

and

temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

seas. Its length is up to 1 m (3.3 ft). Several stranding and mass stranding events have occurred on beaches near

Albany, Western Australia Albany ( ; ) is a port city in the Great Southern region in the Australian state of Western Australia, southeast of Perth, the state capital. The city centre is at the northern edge of Princess Royal Harbour, which is a part of King G ...

. The first South Australian specimen was found at Aldinga in 1944. A cast was made from it, and a replica was made, painted and prepared for display at the South Australian museum that year. Several other individuals have stranded in

South Australia South Australia (commonly abbreviated as SA) is a States and territories of Australia, state in the southern central part of Australia. With a total land area of , it is the fourth-largest of Australia's states and territories by area, which in ...

at Port Willunga,

Netley Netley, officially Netley Abbey, is a village on the south coast of Hampshire, England. It is situated to the south-east of the city of Southampton, and flanked on one side by the ruins of Netley Abbey and on the other by the Royal Victoria Co ...

and West Beach with the latter successfully returned live to deeper water. In contrast to its much larger relatives in the family Molidae, who are very slow-moving and mostly feed upon jellyfish, salps, and small fish and crustaceans, the slender sunfish is known to mainly feed upon squid, particularly of the family

Ommastrephidae Ommastrephidae is a family (biology), family of squid containing three subfamilies, 11 genus, genera, and over 20 species. They are widely distributed globally and are extensively fished for food. One species, ''Todarodes pacificus'', comprised a ...

, which are known for being very fast-moving, displaying evidence that the slender sunfish itself is a faster-moving and agile predator of squid. Several other fossil species of ''Ranzania'' are known: * †'' Ranzania grahami'' Weems, 1985 -

Middle Miocene The Middle Miocene is a sub-epoch of the Miocene epoch (geology), epoch made up of two Stage (stratigraphy), stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0. ...

(

Langhian The Langhian is, in the ICS geologic timescale, an age or stage in the middle Miocene Epoch/Series. It spans the time between 15.97 ± 0.05 Ma and 13.65 ± 0.05 Ma (million years ago) during the Middle Miocene.GeoWhen (2007) The Langhian was ...

Serravallian The Serravallian is, in the geologic timescale, an List of time periods, age or a stage (stratigraphy), stage in the middle Miocene epoch (geology), Epoch/series (stratigraphy), Series, which spans the time between 13.82 annum, Ma and 11.63 Ma (m ...

) of

Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...

, USA * †'' Ranzania ogaii'' Uyeno & Sakamoto, 1994 - Middle Miocene (Langhian) of Saitama, Japan * †'' Ranzania tenneyorum'' Weems, 1985 -

Early Miocene The Early Miocene (also known as Lower Miocene) is a sub-epoch of the Miocene epoch (geology), Epoch made up of two faunal stage, stages: the Aquitanian age, Aquitanian and Burdigalian stages. The sub-epoch lasted from 23.03 ± 0.05 annum, Ma to ...

(

Burdigalian The Burdigalian is, in the geologic timescale, an age (geology), age or stage (stratigraphy), stage in the early Miocene. It spans the time between 20.43 ± 0.05 annum, Ma and 15.97 ± 0.05 Ma (million years ago). Preceded by the Aquitanian (sta ...

) of Virginia, USA * †'' Ranzania zappai'' Carnevale, 2007 ''-'' Middle Miocene (Serravallian) of Italy (named for

Frank Zappa Frank Vincent Zappa (December 21, 1940 – December 4, 1993) was an American guitarist, composer, and bandleader. In a career spanning more than 30 years, Zappa composed Rock music, rock, Pop music, pop, jazz, jazz fusion, orchestra ...

) In addition, a partial fossil of an indeterminate ''Ranzania'' is known from the latest

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

(

Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...

) of

Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...

. An indeterminate

-aged molid, known from a single fossil beak from the Gaiman Formation of

Chubut Chubut may refer to: * Chubut Province Chubut ( from Tehuelche language, Tehuelche 'transparent'; ) is a provinces of Argentina, province in southern Argentina, situated between the 42nd parallel south (the border with Río Negro Province), ...

, Argentina, may also be that of ''Ranzania''; this fossil is notable for representing the southernmost record of fossil

Tetraodontiformes Tetraodontiformes (), also known as the Plectognathi, is an order of ray-finned fishes which includes the pufferfishes and related taxa. This order has been classified as a suborder of the order Perciformes, although recent studies have found ...

. During the warmer climate of the Miocene, the now-frigid coast of Chubut may have been more hospitable for pelagic molids like ''Ranzania''.

Gallery

Image:Sunfish_close_up.jpg, Close-up of the head of a slender sunfish Image:Sunfish_full.jpg, Slender sunfish in Napili Bay, Maui, Hawaii

References

Abu El-Regal and El-Moselhy. 2013. The first record of slender sunfish, Ranzania laevis from the Red Sea. ''Journal of Fish Biology''. 83(5):1425-9 DOI: 10.1111/jfb.12226 slender sunfish Cosmopolitan fish slender sunfish Taxa named by Thomas Pennant {{Tetraodontiformes-stub Extant Burdigalian first appearances