''Silvisaurus'', from the
Latin
Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
silva "woodland" and
Greek
Greek may refer to:
Anything of, from, or related to Greece, a country in Southern Europe:
*Greeks, an ethnic group
*Greek language, a branch of the Indo-European language family
**Proto-Greek language, the assumed last common ancestor of all kno ...
sauros "lizard", is a
nodosaurid
Nodosauridae is a family of ankylosaurian dinosaurs known from the Late Jurassic to the Late Cretaceous periods in what is now Asia, Europe, North America, and possibly South America. While traditionally regarded as a monophyletic clade as the s ...
ankylosaur from the
Early to
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period. It is one of the only known dinosaur species named from
Kansas
Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...
and was designated the official state land fossil in 2023.
Discovery and species

A
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
of the species was discovered in the 1950s by rancher Warren W. Condray of Wells, Kansas. He notified senator
Frank Carlson who directed him to the chancellor of the
University of Kansas
The University of Kansas (KU) is a public research university with its main campus in Lawrence, Kansas, United States. Two branch campuses are in the Kansas City metropolitan area on the Kansas side: the university's medical school and hospital ...
,
Franklin David Murphy. Murphy sent the preparator of the paleontology of vertebrates department of the natural history museum of the university, Russell R. Camp, to investigate the matter. In July 1955 Camp, with help from Condray, recovered the skeleton of a dinosaur. It was further prepared by Camp and Glenn H. Marihugh. In 1960 the find was described and named by
Theodore H. Eaton Jr., also from the University of Kansas, as the
type species
In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
''Silvisaurus condrayi''. The generic name is derived from Latin ''silva'', "wood", in reference to the probably densely forested habitat of the animal. The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
honours Condray.
[Eaton, T.H., Jr. 1960.]
A new armored dinosaur from the Cretaceous of Kansas
. ''The University of Kansas Paleontological Contributions: Vertebrata'' 8: 1-24. To date, ''Silvisaurus'' includes only the type species.
The
holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
, KU 10296, was found in exposures of the
Terra Cotta Clay Member of the
Dakota Formation
The Dakota is a sedimentary rock, sedimentary geologic unit name of Formation (stratigraphy), formation and Group (stratigraphy), group rank in Midwestern North America. The Dakota units are generally composed of sandstones, mudstones, clays, and ...
(late
Albian
The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
-early
Cenomanian
The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) in
Kansas
Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...
, and consists of an incomplete skeleton with skull. It includes the mandible, eight neck vertebrae, ten dorsal vertebrae, a
sacrum
The sacrum (: sacra or sacrums), in human anatomy, is a triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30.
The sacrum situates at the upper, back part of the pelvic cavity, ...
of six sacral vertebrae, three tail vertebrae, a left pubis fragment, the lower end of the right femur, and a toe phalanx. Additionally disarticulated plates and spikes from the body armour were discovered. The condition of the fossil was poor as the bones had been exposed at the bottom of a dry riverbed and had been weathered and trampled by cattle. Some elements were only present as impressions or natural casts.
Description

Based on the remains, the animal is estimated to have been approximately in length. Its skull measures in length and is wide. The bony
secondary palate
The secondary palate is an anatomical structure that divides the nasal cavity from the oral cavity in many vertebrates.
In human embryology, it refers to that portion of the hard palate that is formed by the growth of the two palatine shelves med ...
is poorly developed in ''Silvisaurus'', the
dentary
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone ...
includes at least twenty-five teeth, the basal tubera of the
basioccipital are bulbous, and each
premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammals h ...
holds eight to nine teeth.
The presence of teeth at the front of the jaw suggests that this may have been a relatively primitive nodosaur, since most later forms had a toothless beak instead. In addition to the usual rounded and polygonal
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amph ...
s, ''Silvisaurus'' may have also sported bony spines on its shoulders and tail. The head contained large air passages, which may have been used for loud vocalisations, presumably for communication.
Classification
This taxon represents a relatively primitive nodosaurid, and Vickaryous ''et al.'' (2004) have stated that "''
Sauropelta
''Sauropelta'' ( ; meaning 'lizard shield') is a genus of nodosaurid dinosaur that existed in the Early Cretaceous Period of North America. One species (''S. edwardsorum'') has been named. Anatomically, ''Sauropelta'' is one of the most well-und ...
edwardsorum'', ''Silvisaurus condrayi'', and ''
Pawpawsaurus
''Pawpawsaurus'', meaning "Pawpaw Lizard", is a nodosaurid ankylosaur from the Cretaceous (late Albian) of Tarrant County, Texas, discovered in May 1992. The only species yet assigned to this taxon, ''Pawpawsaurus campbelli,'' is based on a compl ...
campbelli'' form a
basal polytomy
An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree ...
nested deep to ''
Cedarpelta
''Cedarpelta'' is an extinct genus of basal ankylosaurid dinosaur from Utah that lived during the Late Cretaceous period (Cenomanian to lower Turonian stage, 98.2 to 93 Ma) in what is now the Mussentuchit Member of the Cedar Mountain Formation. ...
''."
[Vickaryous, Maryanska, and Weishampel 2004. Chapter Seventeen: Ankylosauria. in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press.]
See also
*
Timeline of ankylosaur research
This timeline of ankylosaur research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ankylosaurs, quadrupedal herbivorous dinosaurs who were protected by a covering bony plates and spik ...
References
Further reading
*Vickaryous,
Maryanska, and Weishampel 2004. Chapter Seventeen: Ankylosauria. in The Dinosauria (2nd edition), Weishampel, D. B., Dodson, P., and Osmólska, H., editors. University of California Press.
{{Taxonbar, from=Q133169
Nodosauridae
Dinosaur genera
Albian dinosaurs
Cenomanian dinosaurs
Fossil taxa described in 1960
Dinosaurs of the United States