Sigmurethra is a
taxonomic category of air-breathing
land snail
A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...
s and
slug
Slug, or land slug, is a common name for any apparently shell-less Terrestrial mollusc, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
s,
terrestrial pulmonate
Pulmonata or pulmonates is an informal group (previously an order, and before that, a subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a pallial lung instead of a gill, or gills. The group inclu ...
gastropod
Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, freshwater, and fro ...
molluscs
Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...
. This is an informal group which includes most land snails and
slug
Slug, or land slug, is a common name for any apparently shell-less Terrestrial mollusc, terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced ...
s.
The two strong synapomorphies of Sigmurethra are a long pedal gland placed beneath a membrane and retractile tentacles.
Several families in this group contain species of snails and slugs that create
love dart
A love dart (also known as a gypsobelum, shooting darts, or just as darts) is a sharp, calcium carbonate, calcareous or chitinous Dart (missile), dart which some hermaphroditic land snails and slugs create. Love darts are both formed and stor ...
s.
Sigmurethra are known from the
Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
to the
Recent periods.
Taxonomy
2005 Taxonomy
In the
taxonomy of the Gastropoda by Bouchet & Rocroi, 2005, Sigmurethra is an "Informal Group", a subsection of the
Stylommatophora.
[Bouchet P. & Rocroi J.-P. (Ed.); Frýda J., Hausdorf B., Ponder W., Valdés Á. & Warén A. 2005. ''Classification and nomenclator of gastropod families''. Malacologia: International Journal of Malacology, 47(1-2). ConchBooks: Hackenheim, Germany. . . 397 pp. http://www.vliz.be/Vmdcdata/imis2/ref.php?refid=78278]
It consists of the following families:
*Superfamily
Clausilioidea
**Family
Clausiliidae
** † Family
Anadromidae
** † Family
Filholiidae
** † Family
Palaeostoidae
*Superfamily
Orthalicoidea
**Family
Orthalicidae
**Family
Cerionidae
**Family
Coelociontidae
** † Family
Grangerellidae
**Family
Megaspiridae
**Family
Placostylidae
**Family
Urocoptidae
*Superfamily
Achatinoidea
**Family
Achatinidae
**Family
Ferussaciidae
**Family
Micractaeonidae
**Family
Subulinidae
*Superfamily
Aillyoidea
**Family
Aillyidae
*Superfamily
Testacelloidea
**Family
Testacellidae
**Family
Oleacinidae
**Family
Spiraxidae
*Superfamily
Papillodermatoidea
**Family
Papillodermatidae
*Superfamily
Streptaxoidea
**Family
Streptaxidae
*Superfamily
Rhytidoidea
**Family
Rhytididae
**Family
Chlamydephoridae
''Chlamydephorus'' is a genus of air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the family Chlamydephoridae. It is the only genus within the family Chlamydephoridae.Herbert D. G. (1997). "The terrestrial slugs of KwaZul ...
**Family
Haplotrematidae
**Family
Scolodontidae
*Superfamily
Acavoidea
**Family
Acavidae
**Family
Caryodidae
**Family
Dorcasiidae
**Family
Macrocyclidae
**Family
Megomphicidae
**Family
Strophocheilidae
Strophocheilidae is a Taxonomy (biology), taxonomic family (biology), family of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Acavoidea (according to the Taxonomy of the Gastropoda (B ...
*Superfamily
Plectopyloidea
**Family
Plectopylidae
**Family
Corillidae
**Family
Sculptariidae
*Superfamily
Punctoidea
**Family
Punctidae
** † Family
Anostomopsidae
**Family
Charopidae
Charopidae is a Taxonomy (biology), taxonomic family (biology), family of small air-breathing land snails (and semi-slugs such as ''Otoconcha dimidiata''), terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Punctoi ...
**Family
Cystopeltidae
**Family
Discidae
**Family
Endodontidae
**Family
Helicodiscidae
**Family
Oreohelicidae
**Family
Thyrophorellidae
*Superfamily
Sagdoidea
**Family
Sagdidae
limacoid clade
*Superfamily
Staffordioidea
**Family
Staffordiidae
*Superfamily
Dyakioidea
**Family
Dyakiidae
*Superfamily
Gastrodontoidea
**Family
Gastrodontidae
**Family
Chronidae
**Family
Euconulidae
**Family
Oxychilidae
**Family
Pristilomatidae
**Family
Trochomorphidae
**Fossil taxa probably belonging to the
Gastrodontoidea
***Subfamily †
Archaeozonitinae
***Subfamily †
Grandipatulinae
***Subfamily †
Palaeoxestininae
*Superfamily
Parmacelloidea
**Family
Parmacellidae
**Family
Milacidae
**Family
Trigonochlamydidae
*Superfamily
Zonitoidea
**Family
Zonitidae
*Superfamily
Helicarionoidea
**Family
Helicarionidae
**Family
Ariophantidae
**Family
Urocyclidae
*Superfamily
Limacoidea
**Family
Limacidae
**Family
Agriolimacidae
**Family
Boettgerillidae
**Family
Vitrinidae
Informal group Sigmurethra continued
Two superfamilies belongs to clade Sigmurethra, but they are not in the limacoid clade.
*Superfamily
Arionoidea
**Family
Arionidae
**Family
Anadenidae
**Family
Ariolimacidae
**Family
Binneyidae
**Family
Oopeltidae
**Family
Philomycidae
*Superfamily
Helicoidea
Helicoidea is a Taxonomy (biology), taxonomic Taxonomic rank, superfamily of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the order (taxonomy), order Stylommatophora.
Taxonomy
2017 taxonomy and l ...
**Family
Helicidae
Helicidae is a large, diverse family of western Palaearctic, medium to large-sized, air-breathing land snails, sometimes called the "typical snails." It includes some of the largest European land snails, several species are common in anthropoge ...
**Family
Bradybaenidae
**Family
Camaenidae
Camaenidae is a Family (biology), family of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea, the typical snails and their allies. This is one of the most diverse families i ...
**Family
Cepolidae
**Family
Cochlicellidae
**Family
Elonidae
**Family
Epiphragmophoridae
**Family
Halolimnohelicidae
**Family
Helicodontidae
**Family
Helminthoglyptidae
**Family
Humboldtianidae
**Family
Hygromiidae
**Family
Monadeniidae
**Family
Pleurodontidae
**Family
Polygyridae
**Family
Sphincterochilidae
**Family
Thysanophoridae
**Family
Trissexodontidae
**Family
Xanthonychidae
(Families that are exclusively fossil are indicated with a dagger †)
New families after 2005
*Superfamily
Streptaxoidea
**
Diapheridae - In 2010, Sutcharit et al. (2010)
[Sutcharit C., Naggs F., Wade C. M., Fontanilla I. & Panha S. (2010). "The new family Diapheridae, a new species of ''Diaphera'' Albers from Thailand, and the position of the Diapheridae within a molecular phylogeny of the Streptaxoidea (Pulmonata: Stylommatophora)". '' Zoological Journal of the Linnean Society'' 160: 1-16. .] established a new family Diapheridae within the Streptaxoidea.
2017 Taxonomy
After excluding groups not related, the informal group Sigmurethra becomes suborder
Helicina, with the following nine infraorders and a collection of families with no superfamily:
*Suborder
Helicina ("Non-Achatinoid Clade")
**Infraorder
Arionoidei
**Infraorder
Clausilioidei
**Infraorder
Helicoidei: formed from
Helicoidea
Helicoidea is a Taxonomy (biology), taxonomic Taxonomic rank, superfamily of air-breathing land snails, terrestrial molluscs, terrestrial pulmonate gastropod mollusks in the order (taxonomy), order Stylommatophora.
Taxonomy
2017 taxonomy and l ...
and
Sagdoidea
**Infraorder
Limacoidei: formed from original ''
limacoid clade''
**Infraorder
Oleacinoidei
***
Oleacinoidea: formed from original
Testacelloidea but with the family
Testacellidae excluded;
***
Haplotrematoidea: formed from original
Rhytidoidea and
Haplotrematidae
**Infraorder
Orthalicoidei
**Infraorder
Pupilloidei (
Orthurethra)
**Infraorder
Rhytidoidei: formed by
Rhytidoidea merging with contents from
Acavoidea.
**Infraorder
Succineoidei (
Elasmognatha)
References
{{Taxonbar, from=Q3483567
Stylommatophora
Extant Cretaceous first appearances
Gastropod taxonomy
Obsolete gastropod taxa