Scleromystax Barbatus
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''Scleromystax'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Callichthyidae Callichthyidae is a family of catfishes (order Siluriformes), called armored catfishes due to the two rows of bony plates (or scutes) along the lengths of their bodies. It contains some of the most popular freshwater aquarium fish, such as many ...
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to small
tributaries A tributary, or an ''affluent'', is a stream or river that flows into a larger stream ('' main stem'' or ''"parent"''), river, or a lake. A tributary does not flow directly into a sea or ocean. Tributaries, and the main stem river into which the ...
from several coastal
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
basins draining the southern and southeastern regions in
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. Most of the
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of ''Scleromystax'' are highly
sexually dimorphic Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...
; males have developed
odontode Odontodes, or dermal teeth, are hard structures found on the external surfaces of animals or near internal openings. They consist of a soft pulp surrounded by dentine and covered by a mineralized substance such as enamel, a structure similar to t ...
s inserted in fleshy
papilla Papilla (Latin, 'nipple') or papillae may refer to: In animals * Papilla (fish anatomy), in the mouth of fish * Papilla (worms), small bumps on the surface of certain worms * Basilar papilla, a sensory organ of lizards, amphibians and fish * ...
e on the preopercular-opercular region and the
dorsal Dorsal (from Latin ''dorsum'' ‘back’) may refer to: * Dorsal (anatomy), an anatomical term of location referring to the back or upper side of an organism or parts of an organism * Dorsal, positioned on top of an aircraft's fuselage The fus ...
and
pectoral fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish aquatic locomotion, swim. Apart from the tail or caudal fin, fish fins have no direct connection with the vertebral column ...
s are 2–3 times as long as those of females. ''S. salmacis'' is an exception, as its sexually dimorphic features are subtle and non-remarkable.


Taxonomy

The species of ''Scleromystax'' were previously classified within the genus ''
Corydoras ''Corydoras'' is a genus of freshwater catfish in the family Callichthyidae and subfamily Corydoradinae. The species usually have more restricted areas of endemism than other callichthyids, but the area of distribution of the entire genus almos ...
''. However, ''Scleromystax'' species are now thought to be more closely related to ''
Aspidoras ''Aspidoras'' is a genus of catfishes of the family Callichthyidae from Brazil. Taxonomy The type species for this genus is '' Aspidoras rochai''. The name ''Aspidoras'' is derived from the Greek ''aspis'' (shield) and ''dora'' (skin). ''Aspido ...
'' in a tribe called Aspidoradini.Britto, M.R. (2003)
Phylogeny of the subfamily Corydoradinae Hoedeman, 1952 (Siluriformes: Callichthyidae), with a definition of its genera.
''Proceedings of the Academy of Natural Sciences of Philadelphia, 153 (1): 119–154.''
Although the
monophyly In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent comm ...
of ''Scleromystax'' has been demonstrated,
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
relationships of its species remains obscure. In addition, the taxonomic status of its species are only partially resolved.Britto, M.R. & Reis, R.E. (2005)
A new ''Scleromystax'' species (Siluriformes: Callichthyidae) from coastal rivers of Southern Brazil.
''Neotropical Ichthyology, 3 (4): 481–488.''


Species

There are currently 5 recognized species in this genus: * '' Scleromystax barbatus'' ( Quoy &
Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon and naturalist. Biography Gaimard was born at Saint-Zacharie on January 31, 1793. He studied medicine at the naval medical school in Toulon, subsequent ...
, 1824)
* '' Scleromystax macropterus'' ( Regan, 1913) * '' Scleromystax prionotos'' ( Nijssen & Isbrücker, 1980) * '' Scleromystax reisi'' M. R. Britto, Fukakusa & L. R. Malabarba, 2016 Britto, M.R., Fukakusa, C.K. & Malabarba, L.R. (2016)
New species of ''Scleromystax'' Günther, 1864 (Siluriformes: Callichthyidae) - extending the meridional distribution of genera endemic to the Atlantic Forest.
''Neotropical Ichthyology, 14 (3): e150158.''
* '' Scleromystax salmacis'' M. R. Britto & R. E. dos Reis, 2005


References

{{Taxonbar, from=Q1756368 Callichthyidae Fish of South America Taxa named by Albert Günther Catfish genera Freshwater fish genera