''Scalamagnus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus of

polycotylid Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widesprea ...

plesiosaur from the

Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...

Tropic Shale Formation The Tropic Shale is a Mesozoic geologic formation. Dinosaur remains are among the fossils that have been recovered from the formation,Weishampel, ''et al.'' (2004). "Dinosaur distribution." Pp. 517-607. including '' Nothronychus graffami''. The ...

of the United States. The genus contains a single species ''S. tropicensis'', known from a skull and two partial skeletons. ''Scalamagnus'' was historically considered to represent a species of the genus ''

Dolichorhynchops ''Dolichorhynchops'' is an extinct genus of polycotylid plesiosaur from the Late Cretaceous of North America, containing the species ''D. osborni'' and ''D. herschelensis'', with two previous species having been assigned to new genera. Definitiv ...

'' before it was moved to its own genus.

History

The taxon was first named as a species of ''

'' by Rebecca Schmeisser McKean in

2012 2012 was designated as: *International Year of Cooperatives *International Year of Sustainable Energy for All Events January *January 4 – The Cicada 3301 internet hunt begins. * January 12 – Peaceful protests begin in the R ...

. The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

is derived from the name of the

Tropic Shale The Tropic Shale is a Mesozoic geologic Formation (geology), formation. Dinosaur remains are among the fossils that have been recovered from the formation,Weishampel, ''et al.'' (2004). "Dinosaur distribution." Pp. 517-607. including ''Nothronych ...

, in which the two specimens were found. It is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

MNA V10046, an almost complete, well-preserved long

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

including most of the

skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...

and from the referred specimen MNA V9431, fragmentary

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

l elements. It was collected between 2003 and 2005 by the

Museum of Northern Arizona The Museum of Northern Arizona is a museum in Flagstaff, Arizona, United States, established as a repository for Indigenous material and natural history specimens from the Colorado Plateau. The museum was founded in 1928 by zoologist Dr. Harol ...

from a single locality within the Tropic Shale of

Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...

, dating to the early

Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...

stage of the early

, about 93.5-91

million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...

. ''D. tropicensis'' extended the known stratigraphic range for ''Dolichorhynchops'' back by approximately 7 million years. Previously three additional polycotylid taxa, '' Eopolycotylus'', '' Palmulasaurus'' and '' Trinacromerum'', have been named from the same formation, two of which are currently

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

to the

. A 2023 study assigns ''D. tropicensis'' to a new genus, ''Scalamagnus''. The generic name is a combination of the

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

''scala'', "staircase", and ''magnus'', "large", referring to the

Grand Staircase-Escalante National Monument Grand may refer to: People with the name * Grand (surname) * Grand L. Bush (born 1955), American actor Places * Grand, Oklahoma, USA * Grand, Vosges, village and commune in France with Gallo-Roman amphitheatre * Grand County (disambiguation), ...

Classification

Clark, O'Keefe & Slack (2023) recovered ''Scalamagnus'' as a

member of the plesiosaur clade

Leptocleidia Plesiosauroidea (; Greek: 'near, close to' and 'lizard') is an extinct clade of carnivorous marine reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroids are known from the Jurassic and Cretaceous perio ...

, as the

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to the clade formed by '' Trinacromerum'' and the

Dolichorhynchia Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widesprea ...

within the

Polycotylinae Polycotylidae is a family of plesiosaurs from the Cretaceous, a sister group to Leptocleididae. They are known as false pliosaurs. Polycotylids first appeared during the Albian stage of the Early Cretaceous, before becoming abundant and widesprea ...

. The results of their

phylogenetic analyses In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as Computational phylogenetics, phylogenetic inference. It infers the relationship among organ ...

are shown in the

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below:

Paleobiology

The holotype is associated with 289

gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In ...

s, which is unusual in comparison to most polycotylid skeletons that generally lack gastroliths. Ranging from less than 0.1 grams to 18.5 grams, the total mass of the gastroliths was about 518 grams. About three-quarters of the stones weighed less than 2 grams, with the mean mass and median mass of the stones respectively estimated at 1.9 grams and 0.8 grams. The gastroliths had high mean value and variability in sphericity, suggesting that this individual was obtaining its stones from rivers located along the western side of the Western Interior Seaway.

References

{{Taxonbar, from=Q123991313 Polycotylidae Late Cretaceous plesiosaurs of North America Sauropterygian genera Fossil taxa described in 2023