The comb duck or American comb duck (''Sarkidiornis sylvicola''), is an unusual

duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...

, found in tropical wetlands in continental

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

south to the

Paraguay River The Paraguay River (''Ysyry Paraguái'' in Guarani language, Guarani, ''Rio Paraguai'' in Portuguese language, Portuguese, ''Río Paraguay'' in Spanish language, Spanish) is a major river in south-central South America, running through Brazil, Bol ...

region in eastern

Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...

, southeastern

Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...

and extreme northeastern

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

, and as a

vagrant Vagrancy is the condition of wandering homelessness without regular employment or income. Vagrants usually live in poverty and support themselves by travelling while engaging in begging, scavenging, or petty theft. In Western countries, ...

Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...

. Most taxonomic authorities split this species and the

knob-billed duck The knob-billed duck (''Sarkidiornis melanotos'') or African comb duck is a type of duck found along the tropical/sub-tropical wetlands and waterways of Sub-Saharan Africa and the island of Madagascar, as well as most of South Asia and mainland I ...

from each other. The comb duck is generally smaller in size when compared to the knob-billed duck, and flanks are darker (black in males, medium grey in females).

Systematics

Uncertainty surrounds the correct

systematic Systematic may refer to: Science * Short for systematic error * Systematic fault * Systematic bias, errors that are introduced by an inaccuracy inherent to the system Economy * Systematic trading, a way of defining trade goals, risk control ...

placement of this species. Initially, it was placed in the

dabbling duck The Anatinae are a subfamily of the family Anatidae (swans, geese and ducks). Its surviving members are the dabbling ducks, which feed mainly at the surface rather than by diving. The other members of the Anatinae are the extinct moa-nalo, a y ...

subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...

Anatinae The Anatinae are a subfamily of the family Anatidae (swans, geese and ducks). Its surviving members are the dabbling ducks, which feed mainly at the surface rather than by diving. The other members of the Anatinae are the extinct moa-nalo, a ...

. Later, it was assigned to the "

perching duck The term perching ducks is used colloquially to mean any species of ducks distinguished by their readiness to perch high in trees. Until the late 20th century, perching ducks meant ''Cairinini'', a tribe of ducks in the duck, goose and swan fam ...

s", a

paraphyletic Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...

assemblage of waterfowl most of which are intermediate between dabbling ducks and

shelduck The shelducks, most species of which are found in the genus ''Tadorna'' (except for the Radjah shelduck, which is now found in its own monotypic genus ''Radjah''), are a group of large birds in the Tadorninae subfamily of the Anatidae, the biolog ...

s. As the "perching ducks" were split up, the comb duck was moved to the

Tadorninae The Tadornini is a biological tribe that includes the shelducks and sheldgeese, which is placed in subfamily Anatinae of family Anatinae, which includes the ducks and most duck-like waterfowl such as the geese and swans. It has been treated a ...

or shelduck

. Analysis of

mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...

sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is cal ...

s of the cytochrome ''b'' and

NADH dehydrogenase NADH dehydrogenase is an enzyme that converts nicotinamide adenine dinucleotide (NAD) from its reduced form (NADH) to its oxidized form (NAD+). Members of the NADH dehydrogenase family and analogues are commonly systematically named using the f ...

subunit 2

gene In biology, the word gene has two meanings. The Mendelian gene is a basic unit of heredity. The molecular gene is a sequence of nucleotides in DNA that is transcribed to produce a functional RNA. There are two types of molecular genes: protei ...

s, however, suggests that it is a quite basal member of the Anatidae, vindicating the earliest placement, but its closest living relatives cannot be resolved to satisfaction without further study.

Description

This common species is unmistakable. It is one of the largest species of duck. Length can range from , wingspan ranges from and weight from .''Sarkidiornis melanotos'' (Comb duck, Knob-billed duck)
. biodiversityexplorer.org Adults have a white head freckled with dark spots, and a pure white neck and underparts. The upperparts are glossy blue-black upperparts, with bluish and greenish

iridescence Iridescence (also known as goniochromism) is the phenomenon of certain surfaces that appear gradually to change colour as the angle of view or the angle of illumination changes. Iridescence is caused by wave interference of light in microstru ...

especially prominent on the secondaries (lower arm feathers). The male is much larger than the female, and has a large black knob on the bill. If seen at a distance, immature comb ducks can also be mistaken for a fulvous whistling duck (''Dendrocygna bicolor''). However, knob-billed ducks in immature

plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...

are rarely seen without adults nearby and thus they are usually easily identified, too.

Ecology

It breeds in still

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...

swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...

s and

lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...

s in the

tropics The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

. It is largely resident, apart from dispersion in the wet season. This

feeds on vegetation by grazing or dabbling and to a lesser extent on small fish,

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s, and seeds. It can become a problem to rice farmers. Knob-billed ducks often perch in trees. They are typically seen in flocks, small in the wet season, up to 100 in the dry season. Sometimes they separate according to sex.

Reproduction

Comb ducks nest mainly in tree holes, also in tall grass. They line their nests with reeds, grass, or feathers, but not down. Males may have two mates at once or up to five in succession. They defend the females and young, but not the nest sites. Unmated males perch in trees and wait for opportunities to mate. Females lay seven to 15 yellowish-white eggs. Several females may lay in a single "dump nest" containing up to 50 eggs.

References

{{Taxonbar, from=Q3473539 Sarkidiornis Taxa named by Hermann von Ihering Taxa named by Rodolpho von Ihering Birds described in 1907