The Santa Maria Formation is a

sedimentary Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles ...

rock formation A rock formation is an isolated, scenic, or spectacular surface rock outcrop. Rock formations are usually the result of weathering and erosion sculpting the existing rock. The term ''rock formation'' can also refer to specific sedime ...

found in Rio Grande do Sul,

Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...

. It is primarily

Carnian The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followe ...

in age (

Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...

), and is notable for its

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

s of

cynodont The cynodonts () ( clade Cynodontia) are a clade of eutheriodont therapsids that first appeared in the Late Permian (approximately 260 mya), and extensively diversified after the Permian–Triassic extinction event. Cynodonts had a wide variet ...

s, " rauisuchian"

pseudosuchia Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to ...

ns, and early

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

s and other dinosauromorphs, including the

herrerasaur Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from dracohors, separate from dinosauria altogether. They are among the oldest known dinosaurs, fi ...

id '' Staurikosaurus'', the

basal Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''. Science * Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure * Basal (medicine), a minimal level that is nec ...

sauropodomorphs Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had l ...

'' Buriolestes'' and ''

Saturnalia Saturnalia is an ancient Roman festival and holiday in honour of the god Saturn, held on 17 December of the Julian calendar and later expanded with festivities through to 23 December. The holiday was celebrated with a sacrifice at the Temple o ...

,'' and the lagerpetid ''

Ixalerpeton ''Ixalerpeton'' (meaning "leaping reptile") is a genus of lagerpetid avemetatarsalian containing one species, ''I. polesinensis''. It lived in the Late Triassic of Brazil alongside the sauropodomorph dinosaur '' Buriolestes''. Discovery and n ...

''. The formation is named after the city of Santa Maria in the central region of Rio Grande do Sul, where outcrops were first studied. The Santa Maria Formation makes up the majority of the Santa Maria Supersequence, which extends through the entire Late Triassic. The Santa Maria Supersequence is divided into four geological

sequences In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called ...

, separated from each other by short unconformities. The first two of these sequences (Pinheiros-Chiniquá and Santa Cruz sequences) lie entirely within the Santa Maria Formation, while the third (the Candelária sequence) is shared with the overlying

Norian The Norian is a division of the Triassic Period. It has the rank of an age ( geochronology) or stage (chronostratigraphy). It lasted from ~227 to million years ago. It was preceded by the Carnian and succeeded by the Rhaetian. Stratigraphic ...

-age

Caturrita Formation The Caturrita Formation is a rock formation found in Rio Grande do Sul, Brazil. Its sediments were deposited in the Paraná Basin. The formation is from the Upper Triassic and forms part of the Santa Maria Supersequence in the upper section of the ...

. The fourth and youngest sequence (the Mata sequence) is equivalent to the

Rhaetian The Rhaetian is the latest age of the Triassic Period (in geochronology) or the uppermost stage of the Triassic System (in chronostratigraphy). It was preceded by the Norian and succeeded by the Hettangian (the lowermost stage or earliest ...

-age Mata Sandstone. The oldest sequence in the formation is the Pinheiros-Chiniquá Sequence (latest Ladinian-earliest Carnian, ~237 Ma), which is biostratigraphically equivalent to the ''

Dinodontosaurus ''Dinodontosaurus'' (meaning "terrible-toothed lizard") is a genus of dicynodont therapsid. It was medium to large dicynodont of the Triassic (with skull up to long) and had a beak corneum. It lived in the Middle Triassic but disappeared in the ...

'' Assemblage Zone. It is followed by the shorter Santa Cruz Sequence (early Carnian-middle Carnian, ~236 Ma), biostratigraphically equivalent to the ''

Santacruzodon ''Santacruzodon'' is an extinct genus of cynodonts which existed in Brazil during the Triassic period. The type species is ''Santacruzodon hopsoni''. Species ''Santacruzodon hopsoni'', is a species that was collected in 1995 in Santa Cruz do Sul ...

'' Assemblage Zone. The final sequence, which is only partially present within the formation, is the Candelária Sequence (middle Carnian-latest Carnian, ~233-228 Ma). The lower portion of this sequence, coinciding with the upper part of the Santa Maria Formation, is equivalent to the ''Hyperodapedon'' Assemblage Zone.The ''Hyperodapedon'' Assemblage Zone is itself subdivided into ''Hyperodapedon'' Acme Zone (most of the zone, where the

rhynchosaur Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaur ...

''Hyperodapedon'' is widely reported) and ''

Exaeretodon ''Exaeretodon'' is an extinct genus of fairly large, low-slung traversodontid cynodonts from the southern parts of Pangea. Four species are known, from various formations. ''E. argentinus'' is from the Carnian-age (Late Triassic) Cancha de Boch ...

'' Zone (restricted to about three known and sampled localities, where rhynchsaurs are almost completely absent, but the traversodontid cynodont ''Exaeretodon'' is widely reported). These subdivisions are also known as Lower and Upper ''Hyperodapedon'' Assemblage Zone, respectively. U-Pb radiometric dating of a locality in the Upper portion of the Santa Maria Formation found an estimated age of 233.23±0.73 million years ago, putting that locality 1.5 million years older than the

Ischigualasto Formation The Ischigualasto Formation is a Late Triassic fossiliferous formation and Lagerstätte in the Ischigualasto-Villa Unión Basin of the southwestern La Rioja Province and northeastern San Juan Province in northwestern Argentina. The formation d ...

and younger than Los

Chañares Formation The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones ...

. The Santa Maria and Ischigualasto formations are approximately equal as having the earliest dinosaur localities.

Vertebrate paleofauna

Most of the information below was included on a revision of the Triassic faunal successions of the Paraná Basin, southern Brazil by Schultz et al.

Vertebrate paleofauna

Temnospondyls

Synapsids

Dicynodonts

Cynognathians

Probainognathians

Reptiles

Pseudosuchians

Avemetatarsalians

Other reptiles

Formations

See also

References