The Ripon barbel (''Labeobarbus altianalis'') is an

East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territories make up Eastern Africa: Due to the historical ...

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fishes, is a class of bony fish. They comprise over 50% of living vertebrate species. The ray-finned fishes are so called because their fins are webs of skin supported by bony or hor ...

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...

. A notably large

barb Barb or the BARBs or ''variation'' may refer to: People * Barb (name), a list of people and fictional characters with the given name or surname * Barb, a term used by fans of Nicki Minaj to refer to themselves * The Barbs, a band Places * Barb, ...

, its maximum recorded

total length Fish measurement is the measuring of individual fish and various parts of their anatomies. These data are used in many areas of ichthyology, including taxonomy and fisheries biology. Overall length * Standard length (SL) is the length of a fish ...

is .

Taxonomy

Like other

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

n "

s", placement of this

in ''

Barbus ''Barbus'' is a genus of ray-finned fish in the family Cyprinidae. The type species of ''Barbus'' is the common barbel, first described as ''Cyprinus barbus'' and now named ''Barbus barbus''. ''Barbus'' is the namesake genus of the subfamily B ...

'' – the genus of the typical

barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprinid ...

s and relatives – was provisional. Though called "barbel", it is probably not closely enough related to the typical

s – the core group of ''Barbus'' – to be considered congeneric. Several supposedly distinct species have been merged into ''B. altianalis'', and numerous

subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...

have been proposed. None of these are deemed valid. Some authorities place this species in the genus

Labeobarbus ''Labeobarbus'' is a mid-sized ray-finned fish genus in the family Cyprinidae. Its species are widely distributed throughout eastern Africa and especially southern Africa, but also in Lake Tana in Ethiopia. A common name, in particular for th ...

. On the other hand, '' B. longifilis'', '' B. paucisquamatus'' and '' B. somereni'' were once considered subspecies of ''L: altianalis'', the second as sspp. ''lobogenysoides'' and ''paucisquamata'' and the third as ssp. ''urundensis''. Significant

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...

s of ''L. altianalis'' are: * ''Labeobarbus altianalis altianalis'' Boulenger, 1900 * ''Labeobarbus altianalis eduardianus'' Boulenger, 1901 * ''Labeobarbus altianalis labiosa'' Pellegrin, 1933 * ''Labeobarbus altianalis radcliffi'' (''

lapsus In philology, a lapsus (Latin for "lapse, slip, error") is an involuntary mistake made while writing or speaking. Investigations In 1895 an investigation into verbal slips was undertaken by a philologist and a psychologist, Rudolf Meringer and K ...

'') * ''Labeobarbus altianalis radcliffii'' Boulenger, 1903 * ''Labeobarbus bayoni'' Boulenger, 1911 * ''Labeobarbus hollyi'' Lohberger, 1929 * ''Labeobarbus lobogenys'' Boulenger, 1906 * ''Labeobarbus pietschmanni'' Lohberger, 1929 * ''Labeobarbus pietschmanni hollyi'' Lohberger, 1929 * ''Labeobarbus pietschmanni pietschmanni'' Lohberger, 1929 * ''Labeobarbus procatopus'' Boulenger, 1916 * ''Labeobarbus radcliffi'' (''lapsus'') * ''Labeobarbus radcliffii'' Boulenger, 1903 * ''Labeobarbus rueppellii'' Pfeffer, 1896 ''L. procatopus'' was described from the

Amala River Amala may refer to: People * Amala Akkineni, South Indian actress * Amala Chebolu, playback singer in the Telugu film industry, also known as Tollywood * Amala Paul (born 1991), South Indian actress * Amala Shankar (1919–2020), Indian dancer * ...

, which was mistakenly believed to be to the east of

Lake Baringo Lake Baringo is, after Lake Turkana, the most northern of the Kenyan Rift Valley lakes, with a surface area of and an elevation of . The lake is fed by several rivers: the Molo, Perkerra and Ol Arabel. It has no obvious outlet; the waters ar ...

. The Ripon barbel does not occur there, and the

taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

was thus thought to refer to ''

Labeobarbus intermedius ''Labeobarbus intermedius'' is an East African ray-finned fish species in the family Cyprinidae. Like the closely related yellowfish, it is hexaploid. A large species, the maximum recorded standard length is nearly . This species has a subspec ...

''. But the Amala is actually a

tributary A tributary, or affluent, is a stream or river that flows into a larger stream or main stem (or parent) river or a lake. A tributary does not flow directly into a sea or ocean. Tributaries and the main stem river drain the surrounding drainage ...

of the

Mara River The Mara River is a river in that begins in Narok County (Kenya) and in ends in Mara Region (Tanzania), and lies across the migration path of ungulates in the Maasai Mara/Serengeti ecosystem. The River's Flow The Mara River basin covers a sur ...

, where ''L. altianalis'' is found, and ''L. procatopus'' is thus a junior synonym of it.

Distribution and ecology

The Ripon barbel inhabits

Lake Victoria Lake Victoria is one of the African Great Lakes. With a surface area of approximately , Lake Victoria is Africa's largest lake by area, the world's largest tropical lake, and the world's second-largest fresh water lake by surface area after ...

and its

drainage basin A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, t ...

and surrounding areas in the

East African Rift The East African Rift (EAR) or East African Rift System (EARS) is an active continental rift zone in East Africa. The EAR began developing around the onset of the Miocene, 22–25 million years ago. In the past it was considered to be part of a ...

, including

Lake Edward Lake Edward (locally Rwitanzigye or Rweru) is one of the smaller African Great Lakes. It is located in the Albertine Rift, the western branch of the East African Rift, on the border between the Democratic Republic of the Congo (DRC) and Uganda, w ...

George George may refer to: People * George (given name) * George (surname) * George (singer), American-Canadian singer George Nozuka, known by the mononym George * George Washington, First President of the United States * George W. Bush, 43rd Presiden ...

, Kyoga and

Kivu Kivu was the name for a large "region" in the Democratic Republic of the Congo under the rule of Mobutu Sese Seko that bordered Lake Kivu. It included three "Sub-Regions" ("Sous-Régions" in French): Nord-Kivu, Sud-Kivu and Maniema, correspondin ...

, and the

Ruzizi River The Ruzizi (also sometimes spelled Rusizi) is a river, long, that flows from Lake Kivu to Lake Tanganyika in Central Africa, descending from about to about above sea level over its length. The steepest gradients occur over the first , where hy ...

down to its

delta Delta commonly refers to: * Delta (letter) (Δ or δ), a letter of the Greek alphabet * River delta, at a river mouth * D ( NATO phonetic alphabet: "Delta") * Delta Air Lines, US * Delta variant of SARS-CoV-2 that causes COVID-19 Delta may also ...

(though not adjacent

Lake Tanganyika Lake Tanganyika () is an African Great Lake. It is the second-oldest freshwater lake in the world, the second-largest by volume, and the second-deepest, in all cases after Lake Baikal in Siberia. It is the world's longest freshwater lake. ...

) and the upper and middle

Kagera River The Kagera River, also known as Akagera River, or Alexandra Nile, is an East African river, forming part of the upper headwaters of the Nile and carrying water from its most distant source.Stanley, H.M., 1899, Through the Dark Continent, London: G ...

. Records from the Tana River are in error. The countries in which it is found are

Burundi Burundi (, ), officially the Republic of Burundi ( rn, Repuburika y’Uburundi ; Swahili language, Swahili: ''Jamuhuri ya Burundi''; French language, French: ''République du Burundi'' ), is a landlocked country in the Great Rift Valley at the ...

Democratic Republic of the Congo The Democratic Republic of the Congo (french: République démocratique du Congo (RDC), colloquially "La RDC" ), informally Congo-Kinshasa, DR Congo, the DRC, the DROC, or the Congo, and formerly and also colloquially Zaire, is a country in ...

Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...

Rwanda Rwanda (; rw, u Rwanda ), officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of Central Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator ...

Tanzania Tanzania (; ), officially the United Republic of Tanzania ( sw, Jamhuri ya Muungano wa Tanzania), is a country in East Africa within the African Great Lakes region. It borders Uganda to the north; Kenya to the northeast; Comoro Islands and ...

and

Uganda }), is a landlocked country in East Africa East Africa, Eastern Africa, or East of Africa, is the eastern subregion of the African continent. In the United Nations Statistics Division scheme of geographic regions, 10-11-(16*) territor ...

. It lives in diverse

freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. Although the term specifically excludes seawater and brackish water, it does include ...

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

s, from

inshore A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past a ...

waters of lakes to rapidly flowing rivers. This species is an

omnivore An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nutr ...

, eating

water plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...

mollusc Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is esti ...

s and

fish Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of li ...

es. It is fished for

sport Sport pertains to any form of Competition, competitive physical activity or game that aims to use, maintain, or improve physical ability and Skill, skills while providing enjoyment to participants and, in some cases, entertainment to specta ...

and food; its Kagera River population being of some commercial significance. Widespread and apparently rather plentiful for such a large fish due to its

ecological Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...

tolerance and omnivorous habits, the Ripon barbel is not considered a

threatened species Threatened species are any species (including animals, plants and fungi) which are vulnerable to endangerment in the near future. Species that are threatened are sometimes characterised by the population dynamics measure of ''critical depensa ...

by the

IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...

. Local stocks might be temporarily affected by

erosion Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is distin ...

after

deforestation Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...

increasing

turbidity Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality. Fluids can ...

and upsetting the aquatic plant life, and unsustainable growth of the fisheries e.g. along the Kagera must be avoided considering that it is presumably – as per its large size and like many relatives – slow-growing and takes long to mature.

Footnotes

References

* (2007): Evolutionary origin of Lake Tana's (Ethiopia) small ''Barbus'' species: indications of rapid ecological divergence and speciation. '' Anim. Biol.'' 57(1): 39-48. (HTML abstract) * {{Taxonbar, from=Q135100 Labeobarbus Cyprinid fish of Africa Fish of Lake Victoria Taxa named by George Albert Boulenger Fish described in 1900 Taxonomy articles created by Polbot