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Rüppell's warbler (''Curruca ruppeli'') is a typical warbler of the genus ''Curruca''. It breeds in
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
,
Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...
and neighbouring islands. It is migratory, wintering in northeast
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
. This is a rare vagrant to western
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. The name is occasionally cited as "Rueppell's warbler".


Taxonomy and etymology

Together with the Cyprus warbler it forms a
superspecies In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
with dark throats, white malar streaks and light remigial fringes. This in turn is related to the species of Mediterranean and
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
''Sylvia'' warblers that have a naked eye-ring, namely the eastern subalpine warbler, Sardinian warbler and Ménétries's warbler. Both groups have a white malar area, but this may not form a clear streak in the latter group; above the white, the heads of males are uniformly dark. The English name and the specific ''ruppeli'' commemorate the German zoologist and explorer
Eduard Rüppell Wilhelm Peter Eduard Simon Rüppell, also spelled Rueppell (20 November 1794 – 10 December 1884) was a German Natural history, naturalist and List of explorers, explorer, best known for his collections and descriptions of plants and animals from ...
(1794–1884). This is a
monotypic species In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
, meaning there are no recognised
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
.


Description

It is a typical "Curruca" warbler, similar in size but slimmer than the Sardinian warbler. The adults have a plain grey back and paler grey underparts. The bill is fine and pointed, with brown legs and red eyes. The striking male has a black head and, usually, a black throat, separated by a white malar streak ("moustache"). Females have a pale throat, and the head is grey rather than black. Their grey back has a brownish tinge. The song is a slower, deeper rattle than that of the Sardinian warbler.


Ecology


Habitat

It breeds in dry and warm environments, including rocky areas, hillsides covered with scrub and maquis, and from sea level to altitudes of approximately 800–1000 meters. In some regions, such as southeastern Turkey, it has been observed at higher elevations up to 1500 meters, and on Crete, up to 1600 meters. This species occupies a diverse range of dry Mediterranean habitats, from forested areas with sufficient undergrowth to maquis with sparse tree cover. It favors open bushy woods of
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
and
cypress Cypress is a common name for various coniferous trees or shrubs from the ''Cupressus'' genus of the '' Cupressaceae'' family, typically found in temperate climates and subtropical regions of Asia, Europe, and North America. The word ''cypress'' ...
, as well as grassy and rocky terrain with varying densities of scrub cover. Compared to the closely related Sardinian warbler, Rüppell's warbler is less reliant on tall and dense vegetation. During the non-breeding season, it frequents scrubby arid areas, as well as gardens, canebrakes, and hedgerows. On migration, it can be observed on bushy hillsides, mountain slopes, and in desert wadis with
Acacia ''Acacia'', commonly known as wattles or acacias, is a genus of about of shrubs and trees in the subfamily Mimosoideae of the pea family Fabaceae. Initially, it comprised a group of plant species native to Africa, South America, and Austral ...
trees.


Breeding

The nest is a robust cup-shaped structure constructed from grass leaves and stems, often incorporating some vegetable down. The interior is lined with finer materials. Nests are typically situated in dense, often thorny scrub, at heights ranging from 45 to 75 centimeters above the ground. In
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
, egg-laying typically occurs from mid-April to mid-May. Clutches usually consist of four or five eggs.


Diet

This
Insectivore file:Common brown robberfly with prey.jpg, A Asilidae, robber fly eating a hoverfly An insectivore is a carnivore, carnivorous animal or plant which eats insects. An alternative term is entomophage, which can also refer to the Entomophagy ...
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
primarily feeds on adult and
larval A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
insects. However, its diet likely includes other
arthropods Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
as well. During the non-breeding season, it also enriches its diet with berries.


Status

This species has an extensive range and a big population. The population trend is unknown, but it is not believed to be rapid enough to bring it to the threshold of
vulnerability Vulnerability refers to "the quality or state of being exposed to the possibility of being attacked or harmed, either physically or emotionally." The understanding of social and environmental vulnerability, as a methodological approach, involves ...
. For this reasons it is classified as "
least concern A least-concern species is a species that has been evaluated and categorized by the International Union for Conservation of Nature (IUCN) as not being a focus of wildlife conservation because the specific species is still plentiful in the wil ...
" on the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. The European population was estimated at 206,000 to 1,030,000 mature individuals. This represents approximately 95% of the global population, which is preliminarily estimated at 217,000 to 1,080,000 mature individuals.


References

{{Taxonbar, from=Q110257515, from2=Q913438 Rüppell's warbler Birds of Europe Birds of West Asia Wintering birds of Africa Rüppell's warbler