botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

, ramification is the divergence of the

stem Stem or STEM most commonly refers to: * Plant stem, a structural axis of a vascular plant * Stem group * Science, technology, engineering, and mathematics Stem or STEM can also refer to: Language and writing * Word stem, part of a word respon ...

and limbs of a

plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...

into smaller ones, i.e., trunk into

branch A branch, also called a ramus in botany, is a stem that grows off from another stem, or when structures like veins in leaves are divided into smaller veins. History and etymology In Old English, there are numerous words for branch, includ ...

es, branches into increasingly smaller branches, and so on. Gardeners stimulate the process of ramification through

pruning Pruning is the selective removal of certain parts of a plant, such as branches, buds, or roots. It is practiced in horticulture (especially fruit tree pruning), arboriculture, and silviculture. The practice entails the targeted removal of di ...

, thereby making

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

s, and other plants bushier and denser. Short internodes (the section of stem between nodes, i.e., areas where

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are produced) help increase ramification in those plants that form branches at these nodes. Long internodes (which may be the result of over-watering, the over-use of

fertilizer A fertilizer or fertiliser is any material of natural or synthetic origin that is applied to soil or to plant tissues to supply plant nutrients. Fertilizers may be distinct from liming materials or other non-nutrient soil amendments. Man ...

, or a seasonal "growth spurt") decrease a gardener's ability to induce ramification in a plant. A high degree of ramification is essential for the creation of topiary as it enables the topiary artist to carve a bush or

hedge A hedge or hedgerow is a line of closely spaced (3 feet or closer) shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate ...

into a shape with an even surface. Ramification is also essential to practitioners of the art of bonsai as it helps re-create the form and habit of a full-size tree in a small tree grown in a container. The pruning practices of

coppicing Coppicing is the traditional method in woodland management of cutting down a tree to a tree stump, stump, which in many species encourages new Shoot (botany), shoots to grow from the stump or roots, thus ultimately regrowing the tree. A forest ...

and

pollarding Pollarding is a pruning system involving the removal of the upper branches of a tree, which promotes the growth of a dense head of foliage and branches. In ancient Rome, Propertius mentioned pollarding during the 1st century BCE. The practice h ...

induce ramification by removing most of a tree's mass above the root. Fruit tree pruning increases the yield of

orchards An orchard is an intentional plantation of trees or shrubs that is maintained for food production. Orchards comprise fruit- or nut-producing trees that are generally grown for commercial production. Orchards are also sometimes a feature of lar ...

by inducing ramification and thereby creating many vigorous, fruitful branches in the place of a few less-fruitful ones.

External links

Annals of Botany 91: 559-569, 2003

Plant physiology Horticulture Plant anatomy Plant morphology {{plant-physiology-stub