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The hard clam (''Mercenaria mercenaria''), also known as the round clam, hard-shell (or hard-shelled) clam, or the quahog, is an edible marine
bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...
mollusk Mollusca is a phylum of protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000  extant species of molluscs are recognized, making it the second-largest animal phylum after Arthropoda. The ...
that is native to the eastern shores of
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
and
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
from
Prince Edward Island Prince Edward Island is an island Provinces and territories of Canada, province of Canada. While it is the smallest province by land area and population, it is the most densely populated. The island has several nicknames: "Garden of the Gulf", ...
to the
Yucatán Peninsula The Yucatán Peninsula ( , ; ) is a large peninsula in southeast Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north and west of the peninsula from the C ...
. It is one of many unrelated edible bivalves that in the United States are frequently referred to simply as
clam Clam is a common name for several kinds of bivalve mollusc. The word is often applied only to those that are deemed edible and live as infauna, spending most of their lives halfway buried in the sand of the sea floor or riverbeds. Clams h ...
s. Older literature sources may use the systematic name ''Venus mercenaria''; this species is in the family
Veneridae The Veneridae or venerids, common name: Venus (mythology), Venus clams, are a very large family of minute to large, saltwater clams, marine bivalve molluscs. Over 500 living species of venerid bivalves are known, most of which are edible, and m ...
, the venus clams. Confusingly, the "ocean quahog" is a different species, ''
Arctica islandica The ocean quahog (''Arctica islandica'') is a species of edible clam, a marine (ocean), marine bivalve mollusc, mollusk in the family Arcticidae. This species is native to the North Atlantic Ocean, and it is harvested commercially as a food sou ...
'', which, although superficially similar in shape, is in a different family of bivalves: it is rounder than the hard clam, usually has a black
periostracum The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods an ...
, and there is no pallial sinus in the interior of the shell.


Alternative names

The hard clam has many alternative
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...
s. It is also known as the Northern quahog, round clam, or chowder clam. In fish markets, there are specialist names for different sizes of this species of clam. The smallest legally harvestable clams are called countnecks or peanuts, next size up are littlenecks, then topnecks. Above that are the cherrystones, and the largest are called quahogs or chowder clams. Rice, M.A. (1992). The Northern Quahog: Biology of ''Mercenaria mercenaria''. Rhode Island Sea Grant Publication No. RIU-B-92-001, University of Rhode Island, Narragansett. 60 pp.
web link
.
The most distinctive of these names is ''quahog'' ( , , or , also spelt ''quahaug'', ''quohog'' or ''cohog''). The word comes from the Narragansett word "poquauhock", which is similar in
Wampanoag The Wampanoag, also rendered Wôpanâak, are a Native Americans in the United States, Native American people of the Indigenous peoples of the Northeastern Woodlands, Northeastern Woodlands currently based in southeastern Massachusetts and forme ...
and some other
Algonquian languages The Algonquian languages ( ; also Algonkian) are a family of Indigenous languages of the Americas and most of the languages in the Algic language family are included in the group. The name of the Algonquian language family is distinguished from ...
; it is first attested in North American English in 1794. Native polities on the eastern Atlantic seaboard made valuable beads called
wampum Wampum is a traditional shell bead of the Eastern Woodlands tribes of Native Americans. It includes white shell beads hand-fashioned from the North Atlantic channeled whelk shell and white and purple beads made from the quahog or Western ...
from the shells, especially those colored purple; the species name ''mercenaria'' is related to the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word for commerce. Today people living in coastal New England still use Algonquian words for the clam, as they have done for thousands of years. In many areas where
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. Nelu ...
is important, clam farmers have bred specialized versions of these clams with distinctions needed for them to be distinguished in the marketplace. These are quite similar to common "wild type" ''Mercenaria'' clams, except that their shells bear distinctive markings. These are known as the ''notata'' strain of quahogs, which occur naturally in low numbers wherever quahogs are found.


Distribution

Western
Atlantic Ocean The Atlantic Ocean is the second largest of the world's five borders of the oceans, oceanic divisions, with an area of about . It covers approximately 17% of Earth#Surface, Earth's surface and about 24% of its water surface area. During the ...
: Hard clams are quite common throughout
New England New England is a region consisting of six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York (state), New York to the west and by the ...
, north into
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, and all down the Eastern seaboard of the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
to
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
; but they are particularly abundant between
Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer months. The ...
and
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
, where seeding and harvesting them is an important commercial form of
aquaculture Aquaculture (less commonly spelled aquiculture), also known as aquafarming, is the controlled cultivation ("farming") of aquatic organisms such as fish, crustaceans, mollusks, algae and other organisms of value such as aquatic plants (e.g. Nelu ...
. For example, the species is an important member of the suspension-feeding,
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from the Ancient Greek word (), meaning "the depths". ...
fauna of the lower
Chesapeake Bay The Chesapeake Bay ( ) is the largest estuary in the United States. The bay is located in the Mid-Atlantic (United States), Mid-Atlantic region and is primarily separated from the Atlantic Ocean by the Delmarva Peninsula, including parts of the Ea ...
.
Rhode Island Rhode Island ( ) is a state in the New England region of the Northeastern United States. It borders Connecticut to its west; Massachusetts to its north and east; and the Atlantic Ocean to its south via Rhode Island Sound and Block Is ...
is situated right in the middle of "quahog country" and has supplied a quarter of the U.S.'s total annual commercial quahog catch. The quahog is the official shellfish of the U.S. state of
Rhode Island Rhode Island ( ) is a state in the New England region of the Northeastern United States. It borders Connecticut to its west; Massachusetts to its north and east; and the Atlantic Ocean to its south via Rhode Island Sound and Block Is ...
and is the namesake of the fictional Rhode Island town featured in the animated sitcom
Family Guy ''Family Guy'' is an American animated sitcom created by Seth MacFarlane for the Fox Broadcasting Company. The series premiered on January 31, 1999, following Super Bowl XXXIII, with the rest of the first season airing from April 11, 1999. Th ...
. The species has also been introduced and is farmed on the
Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is bounded by the cont ...
coast of North America and in
Great Britain Great Britain is an island in the North Atlantic Ocean off the north-west coast of continental Europe, consisting of the countries England, Scotland, and Wales. With an area of , it is the largest of the British Isles, the List of European ...
and continental
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. It reproduces sexually by females and males shedding
gametes A gamete ( ) is a haploid cell that fuses with another haploid cell during fertilization in organisms that reproduce sexually. Gametes are an organism's reproductive cells, also referred to as sex cells. The name gamete was introduced by the Ge ...
into the water.


Habitat

Shallow water: found intertidally on sand flats, at low tide. Hard Clams are found spread throughout the South Atlantic Region. They can be found at a depth of 15 meters or more.


Parasite

Quahog parasite unknown (QPX) is a
parasite Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives (at least some of the time) on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted str ...
that affects the hard shell clam ''Mercenaria mercenaria''. While little is known about the disease, research is currently under way in several laboratories. This research is fueled by the need to inform aquaculturists, who suffer financially because of the mortality rates in clams that QPX inflicts and the ensuing years in which runs must be left
fallow Fallow is a farming technique in which arable land is left without sowing for one or more vegetative cycles. The goal of fallowing is to allow the land to recover and store Organic compound, organic matter while retaining moisture and disrupting ...
to clear the disease. It was discovered along the coast of
Cape Cod Cape Cod is a peninsula extending into the Atlantic Ocean from the southeastern corner of Massachusetts, in the northeastern United States. Its historic, maritime character and ample beaches attract heavy tourism during the summer months. The ...
, Massachusetts in 1995. Quahog parasite X (or quahog parasite unknown PX disease of the hard clam ''Mercenaria mercenaria'' is caused by a poorly known protistan parasite. Its DNA sequence analysis places the QPX parasite among the thraustochytrid
stramenopile The stramenopiles, also called heterokonts, are protists distinguished by the presence of stiff tripartite external hairs. In most species, the hairs are attached to flagella, in some they are attached to other areas of the cellular surface, an ...
s. The QPX parasite is unicellular and possesses at least one flagellum, like most other stramenopile organisms. Thraustochytrids are common protists in marine sediments and the water column, but only a few thraustochytrids are known as parasites of marine animals. Although QPX disease was first recorded on the Atlantic coast of Canada in the early 1960s, it did not become a major economic problem until its appearance in cultured clams at Prince Edward Island, Massachusetts in 1992, and Virginia in 1997. Infected clams are characterized by the presence of blisters or pustules in the mantle and later by gaping and death.


Human use

In coastal areas of the New England states, Long Island, and New Jersey, restaurants known as raw bars or clam bars specialize in serving littlenecks and topnecks raw on an opened half-shell, usually with a cocktail sauce with
horseradish Horseradish (''Armoracia rusticana'', syn. ''Cochlearia armoracia'') is a perennial plant of the family Brassicaceae (which also includes Mustard plant, mustard, wasabi, broccoli, cabbage, and radish). It is a root vegetable, cultivated and us ...
, and often with
lemon The lemon (''Citrus'' × ''limon'') is a species of small evergreen tree in the ''Citrus'' genus of the flowering plant family Rutaceae. A true lemon is a hybrid of the citron and the bitter orange. Its origins are uncertain, but some ...
. Sometimes littlenecks are steamed and dipped in butter, though not as commonly as their soft-shelled clam cousin the "steamer". Littlenecks are often found in-the-shell in sauces, soups, stews, and clams casino, or substituted for European varieties such as the cockle in southern European seafood dishes. The largest clams are quahogs or chowders and cherrystones; they have the toughest meat and are used in such dishes as
clam chowder Clam chowder is any of several chowder soups in American cuisine containing clams. In addition to clams, common ingredients include diced potatoes, salt pork, and onions. It is believed that clams were used in chowder because of the relative ...
, clam cakes, and stuffed clams, or are minced and mixed into dishes that use the smaller, more tender clams. Historically, Native Americans used the quahog as a component in
wampum Wampum is a traditional shell bead of the Eastern Woodlands tribes of Native Americans. It includes white shell beads hand-fashioned from the North Atlantic channeled whelk shell and white and purple beads made from the quahog or Western ...
, the shell beads exchanged in the
North American fur trade The North American fur trade is the (typically) historical Fur trade, commercial trade of furs and other goods in North America, beginning in the eastern provinces of French Canada and the northeastern Thirteen Colonies, American colonies (soon- ...
. The Narragansetts used the hard clam for food and ornaments. A population of hard clams exists in Southampton Water in
Hampshire Hampshire (, ; abbreviated to Hants.) is a Ceremonial counties of England, ceremonial county in South East England. It is bordered by Berkshire to the north, Surrey and West Sussex to the east, the Isle of Wight across the Solent to the south, ...
,
England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...
. Originally bred in the warm water outflows at Southampton Power Station for use as eel bait, the population became self-sustaining and can now be found in Southampton Water and has also spread to Portsmouth Harbour and Langstone Harbour.


Clams and red tide

The term " red tide" refers to an accumulation of a toxin, such as saxitoxin, produced by marine algae. Filter-feeding shellfish are affected, such as clams, oysters, and mussels. As they filter microorganisms, clams ingest ''K. brevis'' algae. This algae accumulates in the clams' tissues and is toxic to humans when they are consumed. The toxin affects the human
central nervous system The central nervous system (CNS) is the part of the nervous system consisting primarily of the brain, spinal cord and retina. The CNS is so named because the brain integrates the received information and coordinates and influences the activity o ...
. Eating contaminated shellfish, raw or cooked, can be fatal. Some other kinds of algal blooms make the seawater appear red, but red tide blooms do not always discolor the water, nor are they related to tides.


Notes


References

{{DEFAULTSORT:Hard Clam Clams Veneridae Marine molluscs of North America Molluscs of the Atlantic Ocean Molluscs of the United States Molluscs of Mexico Fauna of the Northeastern United States Fauna of the Southeastern United States Molluscs described in 1758 Taxa named by Carl Linnaeus Symbols of Rhode Island Rhode Island cuisine Native American culture Seafood in Native American cuisine Eversole, A. (1987, August). Special Profiles: Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (South Atlantic) eview of Special Profiles: Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (South Atlantic) Google Books. https://books.google.com/books?hl=en&lr=&id=PvuECX0o7wMC&oi=fnd&pg=PR6&dq=what+do+hard+clams+eat&ots=QsSbK-dY6X&sig=gar5L8ow9qxr9ZmawA1I1xP-tJg#v=onepage&q=what%20do%20hard%20clams%20eat&f=false