Sori of ''Ptisana fraxinea'' ''Ptisana'' is a genus in the

eusporangiate fern Eusporangiate ferns are vascular spore plants, whose sporangia arise from several epidermal cells and not from a single cell as in leptosporangiate ferns. Typically these ferns have reduced root systems and sporangia that produce large amounts o ...

family

Marattiaceae Marattiaceae is the only family of extant (living) ferns in the order Marattiales. In the Pteridophyte Phylogeny Group classification of 2016 (PPG I), Marattiales is the only order in the subclass Marattiidae. The family has six genera and about ...

, comprising species historically treated in the genus '' Marattia''. The establishment of this genus follows the 2008 work by Andrew G. Murdock, which supported recognition of this group on the basis of genetic analysis and morphology. ''Ptisana'' can be distinguished from ''Marattia'' by the presence of distinct sutures at the point of leaflet attachment, deeply cut

synangia A sporangium (; from Late Latin, ) is an enclosure in which spores are formed. It can be composed of a single cell or can be multicellular. Virtually all plants, fungi, and many other lineages form sporangia at some point in their life cy ...

, and the absence of labiate sporangial apertures. The name ''Ptisana'' is derived from the Latin word for

pearl barley Pearl barley, or pearled barley, is barley that has been processed to remove its fibrous outer hull and polished to remove some or all of the bran layer. It is the most common form of barley for human consumption because it cooks faster and i ...

, an allusion to the shape of the synangia. ''Ptisana'' has a palaeotropical distribution, with the westernmost extreme of the range in Ascension Island and extending eastward through tropical Africa, Asia, and Oceania. Ferns in this genus are generally quite large, with fronds often reaching 2-3 meters in length; the one known exception to this is ''Ptisana rolandi-principis'', a dwarf species endemic to the Plateau de Dogny in New Caledonia. The basal chromosome number for ''Ptisana'' is ''2n=78'', whereas the one count for ''Marattia'' in the strict sense is ''2n=80''. The type species for the genus is ''

Ptisana salicina ''Ptisana salicina'', or king fern, is a species of fern native to Norfolk Island, New Zealand and the South Pacific. Large and robust with a distinctive tropical appearance, it has fronds up to 5 metres (16 feet +/-) tall that arise from a sta ...

,'' the type specimen of which was collected on Norfolk Island.

Species

*''Ptisana africana'' Christenh. – East Africa *''Ptisana ambulans'' Murdock & C.W.Chen –

Solomon Islands Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its capit ...

*''Ptisana attenutata'' ( Labill.) Murdock – New Caledonia *''Ptisana boivinii'' (Mett. ex Kuhn) Senterre & Rouhan –

Madagascar Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...

*''Ptisana boninensis'' (Nakai) Yonek. –

Bonin Islands The Bonin Islands, also known as the , are an archipelago of over 30 subtropical and tropical islands, some directly south of Tokyo, Japan and northwest of Guam. The name "Bonin Islands" comes from the Japanese word ''bunin'' (an archaic read ...

*''Ptisana costulisora'' ( Altson) Murdock –

New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...

and nearby islands *''Ptisana decipiens'' Murdock & C.W.Chen –

*''Ptisana fraxinea'' (

Sm. __NOTOC__ Sir James Edward Smith (2 December 1759 – 17 March 1828) was an English botanist and founder of the Linnean Society. Early life and education Smith was born in Norwich in 1759, the son of a wealthy wool merchant. He displayed a ...

) Murdock –

Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

, including

Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by t ...

islands *''Ptisana grandifolia'' (

Copel. Edwin Bingham Copeland (September 30, 1873 – March 16, 1964) was an American botanist and agriculturist. He is known for founding the University of the Philippines College of Agriculture at Los Baños, Laguna and for being one of the America's l ...

) Murdock –

*'' Ptisana howeana'' ( W.R.B. Oliver) Murdock –

Lord Howe Island Lord Howe Island (; formerly Lord Howe's Island) is an irregularly crescent-shaped volcanic remnant in the Tasman Sea between Australia and New Zealand, part of the Australian state of New South Wales. It lies directly east of mainland P ...

*''Ptisana kingii'' (

) Christenh. –

*''Ptisana koordersii'' (

Alderw. Cornelis Rugier Willem Karel van Alderwerelt van Rosenburgh (Java Island, 23 December 1863 – The Hague, 1 March 1936) was a Dutch botanist Botany, also called , plant biology or phytology, is the science of plant life and a branch of biol ...

) Christenh. – Sulawesi *''Ptisana laboudalloniana'' Senterre & I.Fabre –

Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...

*''Ptisana melanesica'' ( Kuhn) Murdock –

and nearby islands *''Ptisana mertensiana'' ( C. Presl) Murdock –

Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, and ...

*''Ptisana novoguineensis'' ( Rosenst.) Murdock –

*''Ptisana obesa'' (

Christ Jesus, likely from he, יֵשׁוּעַ, translit=Yēšūaʿ, label=Hebrew/Aramaic ( AD 30 or 33), also referred to as Jesus Christ or Jesus of Nazareth (among other names and titles), was a first-century Jewish preacher and religious ...

) Murdock –

*''Ptisana odontosora'' (

) Senterre & Rouhan – Guinea *''Ptisana oreades'' ( Domin) Murdock – northern Australia *''Ptisana papuana'' (

) Murdock & C.W.Chen –

*''Ptisana pellucida'' ( C. Presl) Murdock –

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

and the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

*''Ptisana platybasis'' (

) Murdock –

*''

Ptisana purpurascens ''Ptisana purpurascens'' is a large fern belonging to the botanical family Marattiaceae. It has a globular rhizome with stipule-like fleshy outgrowths. The leaves are dark green, twice pinnate and up to 1 metre long. Every pinnule has up to six ...

'' ( de Vriese) Murdock – Ascension Island *''Ptisana rigida'' (

) Murdock –

, Sulawesi *''Ptisana robusta'' ( Altson) Senterre & I.Fabre –

São Tomé São Tomé is the capital and largest city of the Central African island country of São Tomé and Príncipe. Its name is Portuguese for " Saint Thomas". Founded in the 15th century, it is one of Africa's oldest colonial cities. History Álvar ...

*''Ptisana rolandi-principis'' (Rosenst.) Christenh. – New Caledonia *''Ptisana salicifolia'' (Schrad.) Senterre & Rouhan –

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. It is bounded to the south by of coastline that stretch along the Atlantic Ocean, South Atlantic and Indian Oceans; to the ...

Zimbabwe Zimbabwe (), officially the Republic of Zimbabwe, is a landlocked country located in Southeast Africa, between the Zambezi and Limpopo Rivers, bordered by South Africa to the south, Botswana to the south-west, Zambia to the north, and ...

Malawi Malawi (; or aláwi Tumbuka: ''Malaŵi''), officially the Republic of Malawi, is a landlocked country in Southeastern Africa that was formerly known as Nyasaland. It is bordered by Zambia to the west, Tanzania to the north and northeas ...

and

Mozambique Mozambique (), officially the Republic of Mozambique ( pt, Moçambique or , ; ny, Mozambiki; sw, Msumbiji; ts, Muzambhiki), is a country located in southeastern Africa bordered by the Indian Ocean to the east, Tanzania to the north, Malawi ...

*''

'' (

) Murdock – king fern; South Pacific islands, including

*''Ptisana sambucina'' ( Blume) Murdock –

Malesia Malesia is a biogeographical region straddling the Equator and the boundaries of the Indomalayan and Australasian realms, and also a phytogeographical floristic region in the Paleotropical Kingdom. It has been given different definitions. Th ...

and

Vietnam Vietnam or Viet Nam ( vi, Việt Nam, ), officially the Socialist Republic of Vietnam,., group="n" is a country in Southeast Asia, at the eastern edge of mainland Southeast Asia, with an area of and population of 96 million, making i ...

*''Ptisana senterreana'' Christenh. –

Cameroon Cameroon (; french: Cameroun, ff, Kamerun), officially the Republic of Cameroon (french: République du Cameroun, links=no), is a country in west-central Africa. It is bordered by Nigeria to the west and north; Chad to the northeast; the C ...

Gabon Gabon (; ; snq, Ngabu), officially the Gabonese Republic (french: République gabonaise), is a country on the west coast of Central Africa. Located on the equator, it is bordered by Equatorial Guinea to the northwest, Cameroon to the nort ...

, Equatorial Guinea *''Ptisana smithii'' (

Mett. Georg Heinrich Mettenius (24 November 1823 – 18 August 1866) was a German botanist born in Frankfurt am Main. He was son-in-law to botanist Alexander Braun (1805–1877). In 1845 he received his medical doctorate from the University of Heidelbe ...

ex Kuhn) Murdock –

Vanuatu Vanuatu ( or ; ), officially the Republic of Vanuatu (french: link=no, République de Vanuatu; bi, Ripablik blong Vanuatu), is an island country located in the South Pacific Ocean. The archipelago, which is of volcanic origin, is east of no ...

, Fiji, Samoas,

Tonga Tonga (, ; ), officially the Kingdom of Tonga ( to, Puleʻanga Fakatuʻi ʻo Tonga), is a Polynesian country and archipelago. The country has 171 islands – of which 45 are inhabited. Its total surface area is about , scattered over in ...

*''Ptisana squamosa'' (

) Murdock –

*''Ptisana sylvatica'' ( Blume) Murdock –

*''Ptisana ternatea'' ( de Vriese) Murdock –

Moluccas The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...

, the

*''Ptisana wernerii'' ( Rosenst.) Christenh.

References

Marattiidae Fern genera {{fern-stub