Perennial Rice Regrowth
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A perennial plant or simply perennial is a
plant Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclud ...
that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and
biennials A biennial plant is a flowering plant that, generally in a temperate climate, takes two years to complete its biological life cycle. Life cycle In its first year, the biennal plant undergoes primary growth, during which its vegetative structures ...
. The term is also widely used to distinguish plants with little or no woody growth (secondary growth in girth) from trees and shrubs, which are also technically perennials. Perennialsespecially small
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants th ...
sthat grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their
rootstock A rootstock is part of a plant, often an underground part, from which new above-ground growth can be produced. It could also be described as a stem with a well developed root system, to which a bud from another plant is grafted. It can refer to a ...
or other overwintering structure, are known as
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
perennials. However, depending on the rigours of local climate (temperature, moisture, organic content in the soil, microorganisms), a plant that is a perennial in its native habitat, or in a milder garden, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings, or from divisions. Tomato vines, for example, live several years in their natural tropical/subtropical habitat but are grown as annuals in
temperate In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout t ...
regions because their above-ground biomass doesn't survive the winter. There is also a class of
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
perennials which lack woody stems, such as ''
Bergenia ''Bergenia'' (elephant-eared saxifrage, elephant's ears) is a genus of ten species of flowering plants in the family Saxifragaceae, native to central Asia, from Afghanistan to China and the Himalayan region. Description They are clump-forming, ...
'' which retain a mantle of leaves throughout the year. An intermediate class of plants is known as
subshrub A subshrub (Latin ''suffrutex'') or dwarf shrub is a short shrub, and is a woody plant. Prostrate shrub is a related term. "Subshrub" is often used interchangeably with "bush".Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Der ...
s, which retain a vestigial woody structure in winter, e.g. ''
Penstemon ''Penstemon'' , the beardtongues, is a large genus of roughly 250 species of flowering plants native mostly to the Nearctic, but with a few species also found in the North American portion of the Neotropics. It is the largest genus of flowering ...
''. The symbol for a perennial plant, based on ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genera. It is the first work to consistently apply binomial names and was the ...
'' by
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
, is represented by the symbol: , which is also the
astronomical symbol Astronomical symbols are abstract pictorial symbols used to represent astronomical objects, theoretical constructs and observational events in European astronomy. The earliest forms of these symbols appear in Greek papyrus texts of late antiq ...
for the planet
Jupiter Jupiter is the fifth planet from the Sun and the List of Solar System objects by size, largest in the Solar System. It is a gas giant with a mass more than two and a half times that of all the other planets in the Solar System combined, but ...
.


Life cycle and structure

Perennial plants are most commonly: *Herbaceous - plants that have foliage and stems that die to the ground at the end of the growing season and which show only primary growth. *Evergreen - with persistent foliage without woody stems. *Woody - plants with persistent above ground stems that survive from one growing season to the next, with primary and secondary growth, or growth in width protected by an outer cortex.
The First-Time Gardener: Growing Plants and Flowers: All the Know-How You Need to Plant and Tend Outdoor Areas Using Eco-friendly Methods
'. Quarto Publishing Group USA; 2 February 2021. . p. 18–.
They can be short-lived (only a few years) or long-lived. They include a wide assortment of plant groups from non-flowering plants like
fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except t ...
s and
liverworts The Marchantiophyta () are a division of non-vascular land plants commonly referred to as hepatics or liverworts. Like mosses and hornworts, they have a gametophyte-dominant life cycle, in which cells of the plant carry only a single set of ge ...
to the highly diverse flowering plants like
orchid Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Along with the Asteraceae, they are one of the two largest families of flowering ...
s,
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns an ...
es, and
woody plant A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until sp ...
s. Plants that flower and fruit only once and then die are termed
monocarpic Monocarpic plants are those that flower and set seeds only once, and then die. The term is derived from Greek (''mono'', "single" + ''karpos'', "fruit" or "grain"), and was first used by Alphonse de Candolle. Other terms with the same meaning are ...
or
semelparous Semelparity and iteroparity are two contrasting reproductive strategies available to living organisms. A species is considered semelparous if it is characterized by a single reproductive episode before death, and iteroparous if it is characteri ...
, these species may live for many years before they flower,
The Biology of Reproduction
'. Cambridge University Press; 10 October 2019. . p. 77–.
for example,
century plant ''Agave americana'', common names century plant, maguey, or American aloe, is a species of flowering plant in the family Asparagaceae, native to Mexico and the United States in Texas. It is cultivated worldwide as an ornamental plant, and ha ...
can live for 80 years and grow 30 meters tall before flowering and dying.
Instant Notes in Plant Biology
'. Taylor & Francis; 15 June 2001. . p. 175–.
However, most perennials are
polycarpic Polycarpic plants are those that flower and set seeds many times before dying. A term of identical meaning is pleonanthic and iteroparous. Polycarpic plants are able to reproduce multiple times due to at least some portion of its meristems being ab ...
(or iteroparous), flowering over many seasons in their lifetime.Jill Bailey.
The Facts on File Dictionary of Ecology and the Environment
'. Infobase Publishing; 2004. . p. 132–.
Perennials invest more resources than annuals into roots, crowns, and other structures that allow them to live from one year to the next. They often have a competitive advantage because they can commence their growth and leaf out earlier in the growing season, and can grow taller than annuals, in doing so they can better compete for space and collect more light.John P. Vogel.
Genetics and Genomics of Brachypodium
'. Springer; 17 February 2016. . p. 315–.
Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of
vegetative reproduction Vegetative reproduction (also known as vegetative propagation, vegetative multiplication or cloning) is any form of asexual reproduction occurring in plants in which a new plant grows from a fragment or cutting of the parent plant or spec ...
rather than seeding. These structures include
bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs duri ...
s,
tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...
s, woody crowns,
rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow hori ...
s, turions, woody stems, or
crowns A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
which allows them to survive periods of
dormancy Dormancy is a period in an organism's life cycle when growth, development, and (in animals) physical activity are temporarily stopped. This minimizes metabolic activity and therefore helps an organism to conserve energy. Dormancy tends to be clo ...
over cold or dry seasons; these structures typically store
carbohydrate In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may or ma ...
s which are used once the dormancy period is over and new growth begins. In
climate Climate is the long-term weather pattern in an area, typically averaged over 30 years. More rigorously, it is the mean and variability of meteorological variables over a time spanning from months to millions of years. Some of the meteorologic ...
s that are warm all year long, perennials may grow continuously.Lynden B. Miller.
Parks, Plants, and People: Beautifying the Urban Landscape
'. Norton; 2009. . p. 87–.
Annuals which complete their life cycle in one growing season, in contrast with perennials, produce seeds as the next generation and die; the seeds may survive cold or dry periods or germinate soon after dispersal depending on the climate. Some perennials retain their foliage year-round; these are
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
perennials.
Deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, aft ...
perennials shed all their leaves part of the year,
Forests And Forest Plants - Volume III
'. EOLSS Publications; 24 February 2009. . p. 153–.
they include herbaceous and woody plants; herbaceous plants have stems that lack hard, fibrous growth, while woody plants have stems with buds that survive above ground during dormancy,Tracy DiSabato-Aust.
The Well-Tended Perennial Garden: The Essential Guide to Planting and Pruning Techniques, Third Edition
'. Timber Press; 22 February 2017. . p. 134–.
some perennials are semi-deciduous, meaning they lose some of their leaves in either winter or summer. Deciduous perennials shed their leaves when growing conditions are no longer suitable for photosynthesis, such as when it is too cold or dry. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season.T.T. Kozlowski.
Shedding of Plants Parts
'. Elsevier; 2 December 2012. . p. 88–.
Some perennial plants are protected from
wildfire A wildfire, forest fire, bushfire, wildland fire or rural fire is an unplanned, uncontrolled and unpredictable fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire ...
s because they have underground roots that produce adventitious shoots, bulbs, Crown (botany), crowns, or Underground stem, stems;R. F. Wagle.
Fire, Its Effects on Plant Succession and Wildlife in the Southwest: Some Effects of Fire on Plant Succession and Variability in the Southwest from a Wildlife Management Viewpoint
'. School of Renewable Natural Resources, University of Arizona; 1981. p. 5.
other perennials like trees and shrubs may have thick Bark (botany), cork layers that protect the stems. Herbaceous perennials from temperate and alpine regions of the world can tolerate the cold during winters. Perennial plants may remain dormant for long periods and then recommence growth and reproduction when the environment is more suitable, while most annual plants complete their life cycle during one growing period, and Biennial plant, biennials have two growing periods. The meristem of perennial plants communicates with the hormones produced due to environmental situations (i.e., seasons), reproduction, and stage of development to begin and halt the ability to grow or flower. There is also a distinction between the ability to grow and the actual task of growth. For example, most trees regain the ability to grow during winter but do not initiate physical growth until the spring and summer months. The start of dormancy can be seen in perennials plants through withering flowers, loss of leaves on trees, and halting of reproduction in both flowering and budding plants. Perennials species may produce relatively large seeds that have the advantage of generating larger seedlings that can better compete with other plants. Perennials also produce seeds over many years.


Cultivation

Perennials that are cultivated include: woody plants like fruit trees grown for their edible fruits; shrubs and trees grown as landscaping Ornamental plant, ornamentals; herbaceous food crops like asparagus, rhubarb, Strawberry, strawberries; and subtropical plants not hardy in colder areas such as tomatoes, eggplant, and coleus (which are treated as annuals in colder areas).Jules Janick.
Horticultural Science
'. W. H. Freeman; 15 February 1986 p. 44
Perennials also include plants grown for their flowering and other ornamental value including: bulbs (like tulips, narcissus, and gladiolus); and lawn grass, and other groundcovers, (such as Vinca, periwinkle and dichondra repens, ''Dichondra''). Each type of plant must be separated differently; for example, plants with fibrous root systems like daylilies, Siberian iris or grasses can be pried apart with two garden forks inserted back to back, or cut by knives. However, plants such as bearded irises have a root system of rhizomes; these root systems should be planted with the top of the rhizome just above ground level, with leaves from the following year showing. The point of dividing perennials is to increase the amount of a single breed of plant in your garden. In the United States more than 900 million dollars worth of potted herbaceous perennial plants were sold in 2019.


Benefits in agriculture

Although most of humanity is fed by the re-sowing of the seeds of Annual plant, annual grain crops, (either naturally or by the manual efforts of man), perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion, capture dissolved nitrogen before it can contaminate ground and surface water, and out-compete weeds (reducing the need for herbicides). These potential benefits of perennials have resulted in new attempts to increase the seed yield of perennial species, which could result in the creation of new perennial grain crops.Cox et al. 200

Retrieved on 2008-11-14
Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass. The The Land Institute, Land Institute estimates that profitable, productive perennial grain crops will take at least 25 years to achieve.


Location

Perennial plants dominate many natural ecosystems on land and in fresh water, with only a very few (e.g. ''Zostera'') occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including the trees and shrubs. Perennial plants are usually better long-term competitors, especially under stable, resource-poor conditions. This is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring. Annual plants have an advantage in disturbed environments because of their faster growth and reproduction rates.Stephen B. Monsen.
Proceedings--ecology and Management of Annual Rangelands
'. United States Department of Agriculture, Forest Service, Intermountain Research Station; 1994. p. 342–.


Types

* Examples of
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which ...
perennials include ''Begonia'', banana, and woody plants like Pines, and Holly. * Examples of deciduous perennials include goldenrod, Mentha, mint, and woody plants like maples and Syringa vulgaris, lilacs. * Examples of
monocarpic Monocarpic plants are those that flower and set seeds only once, and then die. The term is derived from Greek (''mono'', "single" + ''karpos'', "fruit" or "grain"), and was first used by Alphonse de Candolle. Other terms with the same meaning are ...
perennials include ''Agave'' and some species of ''Streptocarpus''. * Examples of Woody plant, woody perennials include woody vines, shrubs, and trees such as maple, pine, and apple trees. * Examples of herbaceous perennials used in agriculture include alfalfa, ''Thinopyrum intermedium'', and Red clover.


List of perennials


Perennial flowers

Perennials grown for their decorative flowers include very many species and types. Examples include * Dahlia * Kniphofia * Hollyhock * Lupin


Perennial fruits

The majority of fruit bearing plants are perennial even in temperate climates. Examples include * Apple * Blackcurrant * Blueberry * Blackberry * Ribes, Currant * Grape * Pear * Plum * Raspberries * Strawberry


Perennial herbs

Many herbs are perennial including these examples: * Fennel * Mentha, Mint * Rosemary * Salvia officinalis, Sage * Thyme


Perennial vegetables

Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Examples of some of the more completely perennial vegetables are: * Asparagus * Chives * Globe artichoke * Jerusalem artichoke * Leek * New Zealand spinach * Rhubarb * Sorrel * Rakkyo * Sea kale * Kale * Sweet potato


Aquatic plants

Many aquatic plants are perennial even though many do not have woody tissue. Examples include * Crassula helmsii - New Zealand Stonecrop * Pontederia cordata - Pickerell weed * Stratiotes aloides - Water Soldier which sinks to the bottom of the pond in winter * Utricularia vulgaris - Common bladderwort which produces turions as its overwintering stage


See also

* * * * *


Notes


References


External links


USDA Plant Hardiness Zone Map

Gardening with Perennials



Plants for a Future
{{Authority control Garden plants Perennial plants,