A protoconch (meaning first or earliest or original shell) is an

embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...

nic or

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

l shell which occurs in some classes of

mollusc Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...

s, e.g., the initial chamber of an

ammonite Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...

or the larval shell of a

gastropod Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, freshwater, and fro ...

. In older texts it is also called "nucleus". The protoconch may sometimes consist of several

whorl A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). In nature File:Photograph and axial plane floral diagra ...

s, but when this is the case, the whorls show no growth lines. The whorls of the adult shell, which are formed after the protoconch, are known as the

teleoconch The gastropod shell is part of the body of many gastropods, including snails, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium ...

. The teleoconch starts forming when the larval gastropod becomes a juvenile, and the protoconch may dissolve. Quite often there is a visible line of demarcation where the protoconch ends and the teleoconch begins, and there may be a noticeable change in

sculpture Sculpture is the branch of the visual arts that operates in three dimensions. Sculpture is the three-dimensional art work which is physically presented in the dimensions of height, width and depth. It is one of the plastic arts. Durable sc ...

, or a sudden appearance of sculpture at that point. In some gastropod groups (such as the

Architectonicidae Architectonicidae, common name the staircase shells or sundials, are a family of sea snails, marine gastropod mollusks in the informal group Lower Heterobranchia (= Allogastropoda) of the clade Heterobranchia. The extinct families † Amphit ...

), the teleoconch whorls spiral in the opposite direction to the protoconch. In those cases, the shell is called heterostrophic. In species which have a

veliger A veliger is the planktonic larva of many kinds of sea snails and freshwater snails, as well as most bivalve molluscs (clams) and tusk shells. Description The veliger is the characteristic larva of the gastropod, bivalve and scaphopod taxono ...

or swimming larval stage which hatches out of egg capsules, there are two parts to the protoconch. The first part of the protoconch (which is formed within the embryonic egg capsule) is called protoconch 1, while the part that is formed after the larva has hatched is called protoconch 2. There is often a different sculpture or ornamentation on protoconch 1 compared with protoconch 2, and this can be distinguished under the

microscope A microscope () is a laboratory equipment, laboratory instrument used to examine objects that are too small to be seen by the naked eye. Microscopy is the science of investigating small objects and structures using a microscope. Microscopic ...

. The structure of the protoconch has been widely used as a discriminating feature in gastropod

systematics Systematics is the study of the diversification of living forms, both past and present, and the relationships among living things through time. Relationships are visualized as evolutionary trees (synonyms: phylogenetic trees, phylogenies). Phy ...

. For example, certain

nudibranch Nudibranchs () are a group of soft-bodied marine gastropod molluscs, belonging to the order Nudibranchia, that shed their shells after their larval stage. They are noted for their often extraordinary colours and striking forms, and they have b ...

es have an 'inflated' protoconch, which does not grow but houses the growing soft tissues of the larva, and this distinguishes them from other gastropods. The homologous structure in

bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...

s (clams) is called the

prodissoconch A prodissoconch (meaning first or earliest or original shell) is an embryonic or larval shell which is present in the larva of a bivalve Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda ...

Examples

Comparison of the whole shell and the protoconch of '' Atlanta lesueurii'':

References

{{Gastropod anatomy Mollusc shells