Polihierax Semitorquatus -Serengeti National Park, Tanzania -pair-8
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The African pygmy falcon (''Polihierax semitorquatus'')—or simply pygmy falcon—is a diminutive raptor native to
eastern Eastern or Easterns may refer to: Transportation Airlines *China Eastern Airlines, a current Chinese airline based in Shanghai * Eastern Air, former name of Zambia Skyways *Eastern Air Lines, a defunct American airline that operated from 192 ...
and
southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...
. It is the sole species in its
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
, ''Polihierax''. The pygmy falcon is the smallest bird of prey on the African continent, and among the smallest raptors on earth; only the Asian falconets (of the genus ''
Microhierax The typical falconets, ''Microhierax'', are a bird of prey genus in the family (biology), family Falconidae. They are found in southeast Asia and the smallest members of Falconiformes, averaging about in length and in weight. The smallest membe ...
'') are smaller, weighing roughly 30-40 grams less and measuring about 5 cm shorter from head-to-tail.


Description

Adult pygmy falcons are in length and in weight. These birds are white-bellied, extending up to the face, with males having grey on their backsides, and females have a chestnut back. There are white "eye spots" on the nape. Juveniles have a brown back, duller than adult females, with a rufous wash on the breast. The flight feathers of the wings are spotted black and white (more black above, more white below); the tail is barred black and white. Their flight style is low and undulating. With regards to the pygmy falcon's similarly small size and similar feather colors, they superficially resemble some species of
shrike Shrikes () are passerine birds of the family Laniidae. The family is composed of 34 species in two genera. The family name, and that of the larger genus, '' Lanius'', is derived from the Latin word for "butcher", and some shrikes are also known ...
s. The call is "a high-pitched ''kikiKIK'', repeated" (Kenya) or "a 'chip-chip' and a 'kik-kik-kik-kik'" (southern Africa).


Distribution and habitat

The pygmy falcon inhabits dry bush. The subspecies ''P. s. castanonotus'' occurs from
South Sudan South Sudan (), officially the Republic of South Sudan, is a landlocked country in East Africa. It is bordered on the north by Sudan; on the east by Ethiopia; on the south by the Democratic Republic of the Congo, Uganda and Kenya; and on the ...
to
Somalia Somalia, officially the Federal Republic of Somalia, is the easternmost country in continental Africa. The country is located in the Horn of Africa and is bordered by Ethiopia to the west, Djibouti to the northwest, Kenya to the southwest, th ...
and south to
Uganda Uganda, officially the Republic of Uganda, is a landlocked country in East Africa. It is bordered to the east by Kenya, to the north by South Sudan, to the west by the Democratic Republic of the Congo, to the south-west by Rwanda, and to the ...
and
Tanzania Tanzania, officially the United Republic of Tanzania, is a country in East Africa within the African Great Lakes region. It is bordered by Uganda to the northwest; Kenya to the northeast; the Indian Ocean to the east; Mozambique and Malawi to t ...
; ''P. s. semitorquatus'' occurs from
Angola Angola, officially the Republic of Angola, is a country on the west-Central Africa, central coast of Southern Africa. It is the second-largest Portuguese-speaking world, Portuguese-speaking (Lusophone) country in both total area and List of c ...
to northern
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
. This range is estimated to have an area of 2.7 million km2, and the total pygmy falcon population is estimated to be, at its lowest, 100,000 birds, possibly numbering up to one million at its peak.


Behaviour and ecology

Pygmy falcons show physiological traits that have adaptive value in a region affected by environmental variability associated with the
El Niño–Southern Oscillation El Niño–Southern Oscillation (ENSO) is a global climate phenomenon that emerges from variation in winds and sea surface temperatures over the tropical Pacific Ocean. Those variations have an irregular pattern but do have some semblance of cyc ...
cycle as well varying food source. They regulate their body temperature by thermal buffering provided by
sociable weaver The sociable weaver (''Philetairus socius'') is a species of bird in the weaver family, endemic to Southern Africa. It is the only species in its genus ''Philetairus''. It is found in South Africa, Namibia, and Botswana, but its range is cente ...
colonies and communal roosting by the falcons. This is important because this reduces the energy requirements for rest-phase thermoregulation by occupants.


Food and feeding

Pygmy falcons prey on reptiles and insects, but will occasionally prey on small birds, as well as rodents and appropriately-sized, or juvenile, small mammals. They hunt from a high perch, swooping down and pouncing on their targeted prey. However, some potential prey animals attempt to outsmart the cunning falcons; Kalahari tree skinks have apparently been observed ‘eavesdropping’ on nearby
sociable weaver The sociable weaver (''Philetairus socius'') is a species of bird in the weaver family, endemic to Southern Africa. It is the only species in its genus ''Philetairus''. It is found in South Africa, Namibia, and Botswana, but its range is cente ...
birds, who will loudly warn of any approaching danger, enabling the skinks to run for cover.


Breeding

In Kenya, pygmy falcons nest in
white-headed buffalo weaver The white-headed buffalo weaver or white-faced buffalo-weaver (''Dinemellia dinemelli'') is a species of passerine bird in the family Ploceidae native to East Africa. The buffalo part of its name derives from its habit of following the African bu ...
nests, and the ranges of the two birds coincide. In southern Africa, they are found around red-billed buffalo weaver nests but predominantly nest in the vacant rooms of
sociable weaver The sociable weaver (''Philetairus socius'') is a species of bird in the weaver family, endemic to Southern Africa. It is the only species in its genus ''Philetairus''. It is found in South Africa, Namibia, and Botswana, but its range is cente ...
nests, which are large and multichambered—even if the sociable weavers still have an active colony in the nest. Despite being bird-eaters and bigger than sociable weavers, the pygmy falcons largely leave the latter alone, though they do occasionally catch and eat nestlings and even adults. In addition, falcons are obligate users of
white-headed buffalo weaver The white-headed buffalo weaver or white-faced buffalo-weaver (''Dinemellia dinemelli'') is a species of passerine bird in the family Ploceidae native to East Africa. The buffalo part of its name derives from its habit of following the African bu ...
nests; therefore, they experience all of the benefits these nests provide without the energetic cost of building or maintaining them. Weavers also do not maintain these nest chambers occupied by falcons, increasing the likelihood that these particular chambers will break off the main colony . However, falcons appear to have a preference for chambers away from the center of the colony and with shorter entrance tunnels. Although it is likely that falcons would benefit from selecting chambers with better insulation, they are larger than weavers and as a result may struggle to access chambers through longer entrance tunnel. Pygmy falcons mark their nests by the use of their faecal matter, but the reasons behind this is unknown. It has been suggested that this conspicuous faecal mat (white when fresh and turning pinkish with age) present at the entrance of the chamber could deter snakes and reduce the number of ectoparasites. Faecal matter is also thought to boost the immune system of chicks as well as signal to conspecifics. Faecal mats at the entrance of occupied chambers may serve as a social signal to indicate to other falcons that a colony is occupied. On one occasion a male pygmy falcon was recorded killing another male, and then subsequently took over the dead male's territory and reproduced with the dead male's partner. Breeding occurs between August and February, with a peak in September and December. The maximum and most frequent clutch size was three eggs, these falcons usually only breed once or up to three times per season.


Cooperative breeding

Pygmy falcon territories are occasionally inhabited by groups, where there are more than two adults living together and tending nestlings. There are four potential reasons for this behaviour: defence, co-operative polyandry, delayed dispersal of offspring and cooperation, and thermoregulation (warmth). Corroboration for the last is that in winter African pygmy falcons nest further inside the nest of sociable weavers, where there is better insulation.


Status

Pygmy falcons are not threatened.


References


External links

* Pygmy Falcon
Species text in The Atlas of Southern African Birds


{{Taxonbar, from=Q513174 pygmy falcon Birds of prey of Sub-Saharan Africa Birds of East Africa Birds of Southern Africa pygmy falcon pygmy falcon