zoology Zoology ( , ) is the scientific study of animals. Its studies include the anatomy, structure, embryology, Biological classification, classification, Ethology, habits, and distribution of all animals, both living and extinction, extinct, and ...

and

botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

, a paratype is a specimen of an organism that helps define what the

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

of a

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

and other

taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

actually represents, but it is not the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

(and in botany is also neither an isotype nor a syntype). Often there is more than one paratype. Paratypes are usually held in museum research collections. The exact meaning of the term ''paratype'' when it is used in zoology is not the same as the meaning when it is used in botany. In both cases however, this term is used in conjunction with ''holotype''.

Zoology

zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its formal author, t ...

, a paratype is officially defined as "Each specimen of a type series other than the

.", ''

International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted Convention (norm), convention in zoology that rules the formal scientific name, scientific naming of organisms treated as animals. It is also informally known as the I ...

'' In turn, this definition relies on the definition of a "type series". A type series is the material (specimens of organisms) that was cited in the original publication of the new species or subspecies, and was not excluded from being type material by the author (this exclusion can be implicit, e.g., if an author mentions "paratypes" and then subsequently mentions "other material examined", the latter are not included in the type series), nor referred to as a variant, or only dubiously included in the taxon (e.g., a statement such as "I have before me a specimen which agrees in most respects with the remainder of the type series, though it may yet prove to be distinct" would exclude this specimen from the type series). Thus, in a type series of five specimens, if one is the

, the other four will be paratypes. A paratype may originate from a different locality than the holotype. A paratype cannot become a

lectotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...

, though it is eligible (and often desirable) for designation as a

neotype In biology, a type is a particular specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to anchor or centralizes ...

. The

(ICZN) has not always required a type specimen, but any species or subspecies newly described after the end of 1999 must have a designated holotype or syntypes. A related term is allotype, a term that indicates a specimen that exemplifies the opposite sex of the holotype, and is almost without exception designated in the original description, and, accordingly, part of the type series, and thus a paratype; in such cases, it is functionally no different from any other paratype. It has no nomenclatural standing whatsoever, and although the practice of designating an allotype is recognized by the ICZN, it is not a "

name-bearing type Under the ''International Code of Zoological Nomenclature'' (''Code''), the name-bearing type or onomatophore is the biological type that determines the application of a name. Each animal taxon regulated by the ''Code'' at least potentially has a na ...

" and there are no formal rules controlling how one is designated. Apart from species exhibiting strong

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

, relatively few authors take the trouble to designate such a specimen. It is not uncommon for an allotype to be a member of an entirely different species from the holotype, because of an incorrect association by the original author.

Botany

In botanical nomenclature, a paratype is a specimen cited in the original description that may not have been said to be a type. It is not the holotype nor an isotype (duplicate of the holotype). *If no types were specified, then all specimens cited are syntypes. *If more than one specimen was cited as the type, then they are all syntypes, but specimens cited but not listed as types are paratypes. (Articles 9.5 and 9.6). Like other types, a paratype may be specified for taxa at the rank of

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

or below (Article 7). A paratype may be designated as a lectotype if no holotype, isotype, syntype, or isosyntype (duplicate of a syntype) is extant (Article 9.12).

References

{{reflist Biology terminology Botanical nomenclature Zoological nomenclature

Zoology

Botany

See also

References