Pallas's fish eagle (''Haliaeetus leucoryphus''), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish
sea eagle
A sea eagle or fish eagle (also called erne or ern, mostly in reference to the white-tailed eagle) is any of the birds of prey in the genus ''Haliaeetus'' in the bird of prey family Accipitridae.
Taxonomy and evolution
The genus ''Haliaeetus'' ...
. It breeds in the east
Palearctic
The Palearctic or Palaearctic is the largest of the eight biogeographic realms of the Earth. It stretches across all of Eurasia north of the foothills of the Himalayas, and North Africa.
The realm consists of several bioregions: the Euro-Si ...
in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, China,
India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
,
Nepal
Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne,
सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
,
Bangladesh
Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world, with a population exceeding 165 million pe ...
,
Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
and
Bhutan
Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountai ...
. It is listed as
Endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
on the
IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
.
[ It is partially migratory, with ]Central Asia
Central Asia, also known as Middle Asia, is a region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the former ...
n birds wintering among the southern Asian birds in northern India, and also further west to the Persian Gulf
The Persian Gulf ( fa, خلیج فارس, translit=xalij-e fârs, lit=Gulf of Fars, ), sometimes called the ( ar, اَلْخَلِيْجُ ٱلْعَرَبِيُّ, Al-Khalīj al-ˁArabī), is a mediterranean sea in Western Asia. The bo ...
.
Description
The Pallas's fish eagle has a light sandy-brown hood and a whitish face. The wings are darker brown and the back rufous. The long, slender wings (particularly slender for a sea eagle) are rather dark brown underneath. The tail is black with a wide, distinctive white stripe. Juveniles are overall darker, cooler brown with no band on the tail but with several pale areas on the wing, including the underwing coverts and inner primaries. This results in underwings that have a white band in young fish eagles. It takes until the 4th year or so to obtain adult plumage. Among related species,, it mainly overlaps in range with the quite dissimilar much shorter winged and slightly smaller grey-headed fish eagle and scarcely with the larger, bulkier and much broader winged white-tailed eagle
The white-tailed eagle (''Haliaeetus albicilla'') is a very large species of sea eagle widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which includes other diurnal rapto ...
, which also seldom resembles the coloring of the Pallas's.[ This fairly large species measures in length with a ]wingspan
The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the other wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingspan ...
of . Females are generally reported to weigh , with this sample of nine averaging , and are slightly larger than males at , in ten that weighed an average of . However, in some cases Pallas's fish eagles have been reported to weigh as much as and span as much as much as .[Steele, M. (2017). ''Where in the World are Pallas's Fish Eagles? Migration and Ecology of Haliaeetus leucoryphus in Asia''. University of Arkansas.] Thus, their size falls just slightly under the large northern sea eagles (i.e. bald
Hair loss, also known as alopecia or baldness, refers to a loss of hair from part of the head or body. Typically at least the head is involved. The severity of hair loss can vary from a small area to the entire body. Inflammation or scarr ...
, white-tailed and Steller's) and broadly similar to slightly larger than the sea eagles of more tropical central distribution.[
]
Behaviour and ecology
Diet
Its diet consists primarily of large freshwater fish
Fish are aquatic, craniate, gill-bearing animals that lack limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% ...
. However, Pallas's fish eagle seems to have a broadly opportunistic diet much like better known sea eagles.[ They also regularly prey upon ]bird
Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, especially water bird
A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from sea ...
s. Other prey reportedly can include mammals, including leporid
Leporidae is the family of rabbits and hares, containing over 60 species of extant mammals in all. The Latin word ''Leporidae'' means "those that resemble ''lepus''" (hare). Together with the pikas, the Leporidae constitute the mammalian order ...
s and rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are roden ...
s, frog
A frog is any member of a diverse and largely carnivorous group of short-bodied, tailless amphibians composing the order Anura (ανοὐρά, literally ''without tail'' in Ancient Greek). The oldest fossil "proto-frog" '' Triadobatrachus'' is ...
s, reptiles, including snake
Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more j ...
s and terrapins, insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...
s (i.e. wasp
A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder. Th ...
larvae) and carrion
Carrion () is the decaying flesh of dead animals, including human flesh.
Overview
Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...
.[Orta, J., D. A. Christie, G. M. Kirwan, and C.J. Sharpe (2020). ''Pallas's Fish-Eagle (Haliaeetus leucoryphus)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA.] Their prey in Bangladesh
Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world, with a population exceeding 165 million pe ...
were reported to consist of mrigal carp, rohu, catla, ilish
The ilish (''Tenualosa ilisha'') ( bn, ইলিশ, translit=iliś; also known as the ilishi, hilsa, hilsa herring or hilsa shad, is a species of fish related to the herring, in the family Clupeidae. It is a very popular and sought-after food ...
, '' Chitala chitala'', '' Mystus cavasius'', '' Ompok bimaculatus'' and '' Wallago attu''.[ Known water bird prey can include pheasant-tailed jacana, lesser whistling duck, ferruginous duck, ]common pochard
The common pochard (; ''Aythya ferina'') is a medium-sized diving duck. The scientific name is derived from Greek '' aithuia'', an unidentified seabird mentioned by authors including Hesychius and Aristotle, and Latin ''ferina'', "wild gam ...
and little grebe
The little grebe (''Tachybaptus ruficollis''), also known as dabchick, is a member of the grebe family of water birds. The genus name is from Ancient Greek ''takhus'' "fast" and ''bapto'' "to sink under". The specific ''ruficollis'' is from Lat ...
.[ Pallas's fish eagles are known to attack quite large (i.e. about their own size or somewhat larger) adult water birds including ]greylag geese
The greylag goose or graylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus ''Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A ...
, bar-headed geese
The bar-headed goose (''Anser indicus'') is a goose that breeds in Central Asia in colonies of thousands near mountain lakes and winters in South Asia, as far south as peninsular India. It lays three to eight eggs at a time in a ground nest. It ...
and Demoiselle crane
The demoiselle crane (''Grus virgo'') is a species of crane found in central Eurosiberia, ranging from the Black Sea to Mongolia and North Eastern China. There is also a small breeding population in Turkey. These cranes are migratory birds. Bird ...
by assaulting them on the surface of the water and then flying off with the kill. Since the greylag goose species is slightly heavier than the eagle, this is one of the greatest weight-lifting feats ever recorded for a flying bird. Another case of lifting a great load was recorded at the Yamuna
The Yamuna (Hindustani: ), also spelt Jumna, is the second-largest tributary river of the Ganges by discharge and the longest tributary in India. Originating from the Yamunotri Glacier at a height of about on the southwestern slopes of Ba ...
River in north-central India, where an eagle captured a huge carp
Carp are various species of oily freshwater fish from the family Cyprinidae, a very large group of fish native to Europe and Asia. While carp is consumed in many parts of the world, they are generally considered an invasive species in parts of ...
and flew with the struggling fish very low over the water, before dropping it in response to gunfire. The carp was found to have weighed , which is probably roughly twice the weight of the eagle carrying it.[ How they interact with other raptors is not well understood. However, observations of wintering Pallas's fish eagles in Bharatpur suggest they may dominate several other eagle species, select the highest perches and have the highest daily food intake, including over similarly sized eagles such as ]eastern imperial eagle
The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South ...
s and steppe eagles.
Taxonomy
''Aquila leucorypha'' was the scientific name proposed by Peter Simon Pallas
Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussian zoologist and botanist who worked in Russia between 1767 and 1810.
Life and work
Peter Simon Pallas was born in Berlin, the son ...
in 1771, who first described a Pallas's fish eagle that he encountered during his travels in eastern Siberia
Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part o ...
. In the 19th century, it was also placed in the genera '' Falco'' and ''Haliaeetus'' by different authors.
This species is the hardest-to-place sea-eagle. Among the species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
of its genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
, it has no close living relatives. mtDNA
Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondrion, mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mit ...
cytochrome ''b'' sequence
In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is called ...
data is unable to reliably suggest a phylogenetic
In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
place for it among the sea-eagles. However, some information can be drawn from the molecular data, and especially from morphology and biogeography:
This species retains the ancestral dark eye, bill, and talons of the first sea-eagles, shared with the older tropical
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
lineage. It is peculiar insofar as it has a black band at the end of the tail in adult birds, similar to juvenile Madagascar fish eagles (which look like a smaller, darker version of this bird, but are not very closely related). Its distribution indicates that this species evolved fairly independently of other sea-eagle lineages, but the molecular data tentatively suggests it is possibly closer to the Holarctic
The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical regi ...
species. It diverged from its common ancestor with other species soon after the Holarctic
The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical regi ...
and the tropical lineages split. Dependent on the interpretation of a possible Early Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but ...
sea-eagle fossil from Egypt
Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Med ...
, this happened either at the very start or the end of the Oligocene, somewhere between 34 and 25 mya
Mya may refer to:
Brands and product names
* Mya (program), an intelligent personal assistant created by Motorola
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* Midwest Young Artists, a comprehensive youth music program
Codes
* Burmese ...
.
Distribution and habitat
The Pallas's fish eagle occurs in Russia, Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan, Mongolia, China, India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
, Nepal
Nepal (; ne, नेपाल ), formerly the Federal Democratic Republic of Nepal ( ne,
सङ्घीय लोकतान्त्रिक गणतन्त्र नेपाल ), is a landlocked country in South Asia. It is ma ...
, Bangladesh
Bangladesh (}, ), officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world, with a population exceeding 165 million pe ...
, Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
and Bhutan
Bhutan (; dz, འབྲུག་ཡུལ་, Druk Yul ), officially the Kingdom of Bhutan,), is a landlocked country in South Asia. It is situated in the Eastern Himalayas, between China in the north and India in the south. A mountai ...
.[
]
Conservation
The Pallas's fish eagle is listed as Endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching and inv ...
on the IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biol ...
. The global population is estimated at less than 2,500 individuals. Besides direct persecution, humans contribute to the decline of this species through habitat degradation, pollution
Pollution is the introduction of contaminants into the natural environment that cause adverse change. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the ...
, and draining or overfishing
Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in the ...
lake
A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much lar ...
s. In India, the spread of water hyacinth
''Pontederia crassipes'' (formerly ''Eichhornia crassipes''), commonly known as common water hyacinth is an aquatic plant native to South America, naturalized throughout the world, and often invasive outside its native range.[prey
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...]
difficult for the Pallas's fish eagle.[
Its large range is deceptive, as Pallas's fish eagle is rare and isolated throughout its territory and may not breed in large areas of it.]
References
External links
*
*
{{Taxonbar, from=Q271462
Pallas's fish eagle
Birds of prey of Asia
Birds of Bangladesh
Birds of Myanmar
Pallas's fish eagle