Pallas's fish eagle (''Haliaeetus leucoryphus''), also known as Pallas's sea eagle or band-tailed fish eagle, is a large, brownish

sea eagle A sea eagle or fish eagle (also called erne or ern, mostly in reference to the white-tailed eagle) is any of the birds of prey in the subfamily Haliaeetinae of the bird of prey family Accipitridae. Ten extant species exist, currently described w ...

. It breeds in the east

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

in Kazakhstan, Russia, Tajikistan, Turkmenistan, Uzbekistan, Mongolia, China,

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...

Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...

Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...

and

Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...

. It is listed as

Endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...

on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...

. It is partially migratory, with

Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...

n birds wintering among the southern Asian birds in northern India, and also further west to the

Persian Gulf The Persian Gulf, sometimes called the Arabian Gulf, is a Mediterranean seas, mediterranean sea in West Asia. The body of water is an extension of the Arabian Sea and the larger Indian Ocean located between Iran and the Arabian Peninsula.Un ...

Description

The Pallas's fish eagle has a light sandy-brown hood and a whitish face. The wings are darker brown and the back rufous. The long, slender wings (particularly slender for a sea eagle) are rather dark brown underneath. The tail is black with a wide, distinctive white stripe. Juveniles are overall darker, cooler brown with no band on the tail but with several pale areas on the wing, including the underwing coverts and inner primaries. This results in underwings that have a white band in young fish eagles. It takes until the 4th year or so to obtain adult plumage. Among related species,, it mainly overlaps in range with the quite dissimilar much shorter winged and slightly smaller

grey-headed fish eagle The grey-headed fish eagle (''Icthyophaga ichthyaetus'') is a fish-eating bird of prey from Southeast Asia. It is a large stocky raptor with adults having dark brown upper body, grey head and lighter underbelly and white legs. Juveniles are paler ...

and scarcely with the larger, bulkier and much broader winged

white-tailed eagle The white-tailed eagle (''Haliaeetus albicilla''), sometimes known as the 'sea eagle', is a large bird of prey, widely distributed across temperate Eurasia. Like all eagles, it is a member of the family Accipitridae (or accipitrids) which also ...

, which also seldom resembles the coloring of the Pallas's. This fairly large species measures in length with a

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

of . Females are generally reported to weigh , with this sample of nine averaging , and are slightly larger than males at , in ten that weighed an average of . However, in some cases Pallas's fish eagles have been reported to weigh as much as and span as much as much as .Steele, M. (2017). ''Where in the World are Pallas's Fish Eagles? Migration and Ecology of Haliaeetus leucoryphus in Asia''. University of Arkansas. Thus, their size falls just slightly under the large northern sea eagles (i.e.

bald Hair loss, also known as alopecia or baldness, refers to a loss of hair from part of the head or body. Typically at least the head is involved. The severity of hair loss can vary from a small area to the entire body. Inflammation or scarring ...

, white-tailed and Steller's) and broadly similar to slightly larger than the sea eagles of more tropical central distribution.

Behaviour and ecology

Diet

Its diet consists primarily of large freshwater

fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...

. However, Pallas's fish eagle seems to have a broadly opportunistic diet much like better known sea eagles. They also regularly prey upon

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s, especially

water bird A water bird, alternatively waterbird or aquatic bird, is a bird that lives on or around water. In some definitions, the term ''water bird'' is especially applied to birds in freshwater ecosystems, although others make no distinction from seabi ...

s. Other prey reportedly can include

mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...

s, including

leporid Leporidae () is the family of rabbits and hares, containing over 70 species of extant mammals in all. The family name comes from "Lepus", hare in Latin. Together with the pikas, the Leporidae constitute the mammalian order Lagomorpha. Leporidae ...

s and

rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...

frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...

reptile Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...

s, including

snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...

s and

terrapin Terrapins are a group of several species of small turtle (order Testudines) living in freshwater, fresh or brackish water. Terrapins do not form a taxonomic unit and may not be closely related. Many belong to the families Geoemydidae and Emydid ...

insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...

s (i.e.

wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...

larvae) and

carrion Carrion (), also known as a carcass, is the decaying flesh of dead animals. Overview Carrion is an important food source for large carnivores and omnivores in most ecosystems. Examples of carrion-eaters (or scavengers) include crows, vultures ...

.Orta, J., D. A. Christie, G. M. Kirwan, and C.J. Sharpe (2020). ''Pallas's Fish-Eagle (Haliaeetus leucoryphus)'', version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. Their prey in

were reported to consist of

mrigal carp The mrigal carp (''Cirrhinus cirrhosus''), () also known as the white carp, is a species of ray-finned fish in the carp family. Native to streams and rivers in India, the only surviving wild population is in the Cauvery River, leading to its IU ...

, rohu,

catla Catla (''Labeo catla''; ) also known as the major South Asian carp, is an economically important South Asian freshwater fish in the carp family Cyprinidae. It is native to rivers and lakes in northern India, Bangladesh, Myanmar, Nepal, and Pakis ...

ilish The ilish (''Tenualosa ilisha'') (), also known as the ilishi, hilsa, hilsa herring or hilsa shad, is a species of fish related to the herring, in the family Clupeidae. It is a very popular and sought-after food in the Bengal region, and is t ...

, ''

Chitala chitala ''Chitala chitala'' (Assamese: চিতল ''sitawl'', Bengali: চিতল, ''chitol'') is a knifefish from Bangladesh, India, Nepal, and Pakistan, found in the Brahmaputra, Indus, Ganges and Mahanadi River basins. It is sometimes known as ...

'', '' Mystus cavasius'', ''

Ompok bimaculatus ''Ompok bimaculatus'', known as butter catfish, is a species from the Siluridae, sheatfish Family (biology), family native to Asian countries such as Bangladesh, India, Pakistan, and Sri Lanka, but recently identified in Myanmar. It also indetif ...

'' and ''

Wallago attu ''Wallago attu'', boal, or helicopter catfish, is a species of freshwater catfish of the family Siluridae, native to South and Southeast Asia. ''W. attu'' is found in large rivers and lakes in two geographically disconnected regions (disjunct di ...

''. Known water bird prey can include

pheasant-tailed jacana The pheasant-tailed jacana (''Hydrophasianus chirurgus'') is a Jacanidae, jacana in the Monotypic taxon, monotypic genus ''Hydrophasianus''. Like all other jacanas, they have elongated toes and nails that enable them to walk on floating vegetatio ...

lesser whistling duck The lesser whistling duck (''Dendrocygna javanica''), also known as Indian whistling duck or lesser whistling teal, is a species of whistling duck that breeds in the Indian subcontinent and Southeast Asia. They are nocturnal feeders that during ...

ferruginous duck The ferruginous duck (''Aythya nyroca''), also known as ferruginous pochard, common white-eye or white-eyed pochard, is a medium-sized diving duck from Eurosiberia. The scientific name is derived from the Ancient Greek word, (), an unknown se ...

common pochard The common pochard (; ''Aythya ferina''), known simply as pochard in the United Kingdom, is a medium-sized diving duck in the family Anatidae. It is widespread across the Palearctic. It breeds primarily in the steppe regions of Scandinavia and Si ...

and

little grebe The little grebe (''Tachybaptus ruficollis''), also known as dabchick, is a member of the grebe family of water birds. The genus name is from Ancient Greek ''takhus'' "fast" and ''bapto'' "to sink under". The specific ''ruficollis'' is from Latin ...

. Pallas's fish eagles are known to attack quite large (i.e. about their own size or somewhat larger) adult water birds including

greylag geese The greylag goose (''Anser anser'') is a species of large goose in the waterfowl family Anatidae and the type species of the genus '' Anser''. It has mottled and barred grey and white plumage and an orange beak and pink legs. A large bird, it m ...

, bar-headed geese and

Demoiselle crane The demoiselle crane (''Grus virgo'') is a species of crane found in central Eurosiberia, ranging from the Black Sea to Mongolia and Northeast China. There is also a small breeding population in Turkey. These cranes are migratory birds. Birds f ...

by assaulting them on the surface of the water and then flying off with the kill. Since the greylag goose species is slightly heavier than the eagle, this is one of the greatest weight-lifting feats ever recorded for a flying bird. Another case of lifting a great load was recorded at the

Yamuna The Yamuna (; ) is the second-largest tributary river of the Ganges by discharge and the longest tributary in India. Originating from the Yamunotri Glacier at a height of about on the southwestern slopes of Bandarpunch peaks of the Low ...

River in north-central India, where an eagle captured a huge

carp The term carp (: carp) is a generic common name for numerous species of freshwater fish from the family (biology), family Cyprinidae, a very large clade of ray-finned fish mostly native to Eurasia. While carp are prized game fish, quarries and a ...

and flew with the struggling fish very low over the water, before dropping it in response to gunfire. The carp was found to have weighed , which is probably roughly twice the weight of the eagle carrying it. How they interact with other raptors is not well understood. However, observations of wintering Pallas's fish eagles in Bharatpur suggest they may dominate several other eagle species, select the highest perches and have the highest daily food intake, including over similarly sized eagles such as

eastern imperial eagle The eastern imperial eagle (''Aquila heliaca'') is a large bird of prey that breeds in southeastern Europe and extensively through West and Central Asia. Most populations are migratory and winter in northeastern Africa, the Middle East and South a ...

s and

steppe eagle The steppe eagle (''Aquila nipalensis'') is a large bird of prey. Like all eagles, it belongs to the family Accipitridae. The steppe eagle's well-feathered legs illustrate it to be a member of the subfamily Aquilinae, also known as the "booted ...

Taxonomy

''Aquila leucorypha'' was the

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

proposed by

Peter Simon Pallas Peter Simon Pallas Fellow of the Royal Society, FRS FRSE (22 September 1741 – 8 September 1811) was a Prussia, Prussian zoologist, botanist, Ethnography, ethnographer, Exploration, explorer, Geography, geographer, Geology, geologist, Natura ...

in 1771, who first described a Pallas's fish eagle that he encountered during his travels in eastern

Siberia Siberia ( ; , ) is an extensive geographical region comprising all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has formed a part of the sovereign territory of Russia and its predecessor states ...

. In the 19th century, it was also placed in the genera '' Falco'' and ''Haliaeetus'' by different authors. This species is the hardest-to-place sea-eagle. Among the

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

of its

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

, it has no close living relatives.

mtDNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...

cytochrome ''b''

sequence In mathematics, a sequence is an enumerated collection of objects in which repetitions are allowed and order matters. Like a set, it contains members (also called ''elements'', or ''terms''). The number of elements (possibly infinite) is cal ...

data is unable to reliably suggest a

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

place for it among the sea-eagles. However, some information can be drawn from the molecular data, and especially from

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

and

biogeography Biogeography is the study of the species distribution, distribution of species and ecosystems in geography, geographic space and through evolutionary history of life, geological time. Organisms and biological community (ecology), communities o ...

: This species retains the ancestral dark eye,

bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pl ...

, and

talons A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tars ...

of the first sea-eagles, shared with the older

tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

lineage. It is peculiar insofar as it has a black band at the end of the tail in adult birds, similar to juvenile

Madagascar fish eagle The Madagascar fish eagle (''Icthyophaga vociferoides'') or Madagascar sea-eagle (to distinguish it from the ''Ichthyophaga'' fishing-eagles), is a large bird of prey in the family Accipitridae which also includes many other diurnal raptors such ...

s (which look like a smaller, darker version of this bird, but are not very closely related). Its distribution indicates that this species evolved fairly independently of other sea-eagle lineages, but the molecular data tentatively suggests it is possibly closer to the

Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...

species. It diverged from its common ancestor with other species soon after the

and the tropical lineages split. Dependent on the interpretation of a possible Early

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

sea-eagle fossil from

Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...

, this happened either at the very start or the end of the Oligocene, somewhere between 34 and 25 mya.

Distribution and habitat

The Pallas's fish eagle occurs in Russia, Turkmenistan, Uzbekistan, Tajikistan, Kazakhstan, Mongolia, China,

and

Conservation

The Pallas's fish eagle is listed as

on the

. The global population is estimated at less than 2,500 individuals. Besides direct persecution, humans contribute to the decline of this species through habitat degradation,

pollution Pollution is the introduction of contaminants into the natural environment that cause harm. Pollution can take the form of any substance (solid, liquid, or gas) or energy (such as radioactivity, heat, sound, or light). Pollutants, the component ...

, and draining or

overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing Fish stocks, fish stock), resu ...

lake A lake is often a naturally occurring, relatively large and fixed body of water on or near the Earth's surface. It is localized in a basin or interconnected basins surrounded by dry land. Lakes lie completely on land and are separate from ...

s. In India, the spread of

water hyacinth ''Pontederia crassipes'' (formerly ''Eichhornia crassipes''), commonly known as common water hyacinth, is an aquatic plant native to South America, naturalized throughout the world, and often invasive species, invasive outside its native rang ...

in lakes possibly makes finding

prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...

difficult for the Pallas's fish eagle. Its large

range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...

is deceptive, as Pallas's fish eagle is rare and isolated throughout its territory and may not breed in large areas of it.

References

External links

* * {{Taxonbar, from=Q271462 Pallas's fish eagle Birds of prey of Asia Birds of Bangladesh Birds of Myanmar Pallas's fish eagle Eagles Taxa named by Peter Simon Pallas