Bignoniaceae () is a

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...

s in the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Lamiales The Lamiales (also known as the mint order) are an order of flowering plants in the asterids clade of the Eudicots. Under the APG IV system of flowering plant classification the order consists of 24 families, and includes about 23,810 species ...

commonly known as the bignonias or trumpet vines.Vernon H. Heywood, Richard K. Brummitt, Ole Seberg, and Alastair Culham. ''Flowering Plant Families of the World''. Firefly Books: Ontario, Canada. (2007). . It is not known to which of the other families in the order it is most closely related.Peter F. Stevens (2001 onwards). "Bignoniaceae" At:

Angiosperm Phylogeny Website The Angiosperm Phylogeny Website (or APweb) is a website that presents up-to-date research on the phylogeny and taxonomy of flowering plants (angiosperms) in what is intended to be a user-friendly way. The site is hosted by the Missouri Botanical ...

. At: Botanical Databases At:

Missouri Botanical Garden The Missouri Botanical Garden is a botanical garden located at 4344 Shaw Boulevard in St. Louis, Missouri. It is also known informally as Shaw's Garden for founder and philanthropy, philanthropist Henry Shaw (philanthropist), Henry Shaw. I ...

Website. (see ''External links'' below) Nearly all of the Bignoniaceae are

woody plant A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to Herbaceous plant, herbaceous plants that die back to t ...

s, but a few are subwoody, either as

vine A vine is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas, or runners. The word ''vine'' can also refer to such stems or runners themselves, for instance, when used in wicker work.Jackson; Benjamin; Da ...

s or

subshrub A subshrub (Latin ''suffrutex'') or undershrub is either a small shrub (e.g. prostrate shrubs) or a perennial that is largely herbaceous but slightly woody at the base (e.g. garden pink and florist's chrysanthemum). The term is often interch ...

s. A few more are

herbaceous plant Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...

s of high-

elevation The elevation of a geographic location (geography), ''location'' is its height above or below a fixed reference point, most commonly a reference geoid, a mathematical model of the Earth's sea level as an equipotential gravitational equipotenti ...

montane Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s, in three exclusively herbaceous

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

: ''

Tourrettia ''Tourrettia'' is a monotypic genus of flowering plants belonging to the family Bignoniaceae. It only contains one known species, ''Tourrettia lappacea'' (L'Hér.) Willd. The genus of ''Tourrettia'' has 2 known synonyms, ''Dombeya'' and ''Medica ...

'', '' Argylia'', and ''

Incarvillea ''Incarvillea'' is a genus of about 16 species''Incarvillea''.
Flora of China. ...

''. The family includes many

liana A liana is a long-Plant stem, stemmed Woody plant, woody vine that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the Canopy (biology), canopy in search of direct sunlight. T ...

s, climbing by

tendril In botany, a tendril is a specialized Plant stem, stem, leaf or Petiole (botany), petiole with a thread-like shape used by climbing plants for support and attachment, as well as cellular invasion by parasitic plants such as ''Cuscuta''. There ar ...

s, by twining, or rarely, by

aerial root Aerial roots are Root, roots growing above the ground. They are often Plant development#Adventitious structures, adventitious, i.e. formed from nonroot tissue. They are found in diverse plant species, including epiphytes such as orchids ('' ...

s. The largest

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide use of the term in English is in the discipline of anthropology. The definition is contested, in part due to conflict ...

in the family, called Bignonieae, consists mostly of lianas and is noted for its unique

wood anatomy Wood anatomy is a scientific sub-area of wood science, which examines the variations in xylem anatomical characteristics across trees, shrubs, and herbaceous species to explore inquiries related to plant function, growth, and the environment. Ex ...

.Eberhard Fischer, Inge Theisen, and Lúcia G. Lohmann. 2004. "Bignoniaceae". pages 9-38. In: Klaus Kubitzki (editor) and Joachim W. Kadereit (volume editor). ''The Families and Genera of Vascular Plants'' volume VII. Springer-Verlag: Berlin; Heidelberg, Germany. The family has a nearly

cosmopolitan distribution In biogeography, a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats; most cosmopolitan species are known to be highly adaptable to a range of climatic and en ...

, but is mostly

tropical The tropics are the regions of Earth surrounding the equator, where the sun may shine directly overhead. This contrasts with the temperate or polar regions of Earth, where the Sun can never be directly overhead. This is because of Earth's ax ...

, with a few

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

native Native may refer to: People * '' Jus sanguinis'', nationality by blood * '' Jus soli'', nationality by location of birth * Indigenous peoples, peoples with a set of specific rights based on their historical ties to a particular territory ** Nat ...

to the

temperate zones In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

. Its greatest

diversity Diversity, diversify, or diverse may refer to: Business *Diversity (business), the inclusion of people of different identities (ethnicity, gender, age) in the workforce *Diversity marketing, marketing communication targeting diverse customers * ...

is in northern South America. The family has been covered in some major

floristic {{Short pages monitor ''Did you mean: "Floristics Floristics is the study of plants of geographical regions. It is a branch of phytogeography, which technically makes it a branch of botany, geography, and a subbranch of biogeography. Harvard Un ...

projects, such as

Flora of China The flora of China consists of a diverse range of plant species including over 39,000 vascular plants, 27,000 species of fungi and 3000 species of bryophytes.Wu, Z. Y., P. H. Raven & D. Y. Hong, eds. 2006. Flora of China. Vol. 22 (Poaceae). Sci ...

Flora Malesiana ''Flora Malesiana'' is a multi-volume flora describing the vascular plants of Malesia (the biogeographical region consisting of Indonesia, Malaysia, Singapore, Brunei, the Philippines, and Papua New Guinea), published by the National Herbarium of t ...

, and

Flora Neotropica ''Flora Neotropica'' is a series of monographs published by the New York Botanical Garden Press, and is the official publication of the Organization for Flora Neotropica. It covers the taxonomic treatment of American plants and plant families in ...

. It has not yet been covered in some others, such as

Flora of Australia The flora of Australia comprises a vast assemblage of plant species estimated to over 21,000 vascular and 14,000 non-vascular plants, 250,000 species of fungi and over 3,000 lichens. The flora has strong affinities with the flora of Gondwana, ...

, and

Flora of North America The ''Flora of North America North of Mexico'' (usually referred to as ''FNA'') is a multivolume work describing the native plants and naturalized plants of North America, including the United States, Canada, St. Pierre and Miquelon, and Greenla ...

. Bignoniaceae are most noted for

ornamentals Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

, such as ''

Jacaranda ''Jacaranda'' is a genus of 49 species of flowering plants in the family Bignoniaceae, native to tropical and subtropical regions of the Americas while cultivated around the world. The generic name is also used as the common name. The species ' ...

'', ''

Tabebuia ''Tabebuia'' is a genus of flowering plants in the Family (biology), family Bignoniaceae.Eberhard Fischer, Inge Theisen, and Lúcia G. Lohmann. 2004. "Bignoniaceae". pages 9-38. In: Klaus Kubitzki (editor) and Joachim W. Kadereit (volume editor) ...

'' and ''

Spathodea ''Spathodea'' is a genus in the plant family Bignoniaceae. The single species it contains, ''Spathodea campanulata'', is commonly known as the African tulip tree. The tree grows between tall and is native to tropical dry forests of Africa. It h ...

'', grown for their conspicuous, tubular flowers.George W. Staples and Derral R. Herbst. 2005. "A Tropical Garden Flora" Bishop Museum Press: Honolulu, HI, USA. A great many species are known in cultivation.

Anthony Huxley Anthony Julian Huxley (2 December 1920 – 26 December 1992) was a British botanist and writer. An elected council member of the Royal Horticultural Society, he became its vice president in 1991. He edited '' Amateur Gardening'' from 1967 to 19 ...

, Mark Griffiths, and Margot Levy (1992). ''The New Royal Horticultural Society Dictionary of Gardening''. The Macmillan Press, Limited: London. The Stockton Press: New York. (set). Various other uses have been made of members of this family. Several species were of great importance to the

indigenous peoples There is no generally accepted definition of Indigenous peoples, although in the 21st century the focus has been on self-identification, cultural difference from other groups in a state, a special relationship with their traditional territ ...

of the American tropics. '' Fridericia elegans'', ''

Tanaecium bilabiata ''Tanaecium'' is a genus of flowering plants in the family Bignoniaceae, native to south and Central America. Species *''Tanaecium affinis'' (A.H Gentry) L.G. Lohmann *'' Tanaceium apiculatum'' A.H Gentry *''Tanaecium bilabiatum'' (Sprague) L.G ...

'', and ''

Tanaecium excitosum ''Tanaecium'' is a genus of flowering plants in the family Bignoniaceae, native to south and Central America. Species *''Tanaecium affinis'' (A.H Gentry) L.G. Lohmann *'' Tanaceium apiculatum'' A.H Gentry *''Tanaecium bilabiatum'' (Sprague) L.G ...

'' are

poisonous A poison is any chemical substance that is harmful or lethal to living organisms. The term is used in a wide range of scientific fields and industries, where it is often specifically defined. It may also be applied colloquially or figurati ...

livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...

and have caused severe losses. According to different accounts, the number of species in the family is about 810David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. or about 860. The last

monograph A monograph is generally a long-form work on one (usually scholarly) subject, or one aspect of a subject, typically created by a single author or artist (or, sometimes, by two or more authors). Traditionally it is in written form and published a ...

of the entire family was published in 2004. In that

work Work may refer to: * Work (human activity), intentional activity people perform to support themselves, others, or the community ** Manual labour, physical work done by humans ** House work, housework, or homemaking ** Working animal, an ani ...

, 104 genera were recognized. Since that time,

molecular phylogenetic Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ...

studies Study or studies may refer to: General * Education **Higher education * Clinical trial * Experiment * Field of study * Observational study * Scientific study * Research * Study skills, abilities and approaches applied to learning Other * Study ...

have greatly clarified relationships within the family, and the number of accepted genera is now between 80 and 85.Lúcia G. Lohmann and Carmen U. Ulloa. 2007 onward. Bignoniaceae in iPlants prototype Checklist. (See ''External links'' below). Bignoniaceae Bagh-e-Jinnah

Description

Members of this family are mostly

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

s or

s, sometimes

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

s, and rarely

s or

herbs Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnish (food), garnishing food, for medicinal purposes, or for fragrances. Culinary use typi ...

. Lianas of the tribe Bignonieae have a unique

vascular Vascular can refer to: * blood vessels, the vascular system in animals * vascular tissue Vascular tissue is a complex transporting tissue, formed of more than one cell type, found in vascular plants. The primary components of vascular tissue ...

structure A structure is an arrangement and organization of interrelated elements in a material object or system, or the object or system so organized. Material structures include man-made objects such as buildings and machines and natural objects such as ...

, in which

phloem Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is ...

arms extend downward into the

xylem Xylem is one of the two types of transport tissue (biology), tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts o ...

because certain segments of the

cambium A cambium (: cambiums or cambia), in plants, is a tissue layer that provides partially undifferentiated cells for plant growth. It is found in the area between xylem and phloem. A cambium can also be defined as a cellular plant tissue from whic ...

cease the production of xylem at an early stage of

development Development or developing may refer to: Arts *Development (music), the process by which thematic material is reshaped * Photographic development *Filmmaking, development phase, including finance and budgeting * Development hell, when a proje ...

. The number of these arms is four or a multiple thereof, up to 32. When four, the phloem arms appear as a cross, hence, the

common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often con ...

"cross vine". The phloem in the arms has wider

sieve tubes Sieve elements or sieve tube elements are specialized cells that are important for the function of phloem, which is a highly organized tissue that transports organic compounds made during photosynthesis. Sieve elements are the major conducting cell ...

and less

parenchyma upright=1.6, Lung parenchyma showing damage due to large subpleural bullae. Parenchyma () is the bulk of functional substance in an animal organ such as the brain or lungs, or a structure such as a tumour. In zoology, it is the tissue that ...

than the ordinary phloem. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are petiolate. Leaf arrangement usually is

opposite In lexical semantics, opposites are words lying in an inherently incompatible binary relationship. For example, something that is ''even'' entails that it is not ''odd''. It is referred to as a 'binary' relationship because there are two members i ...

, or rarely

alternate Alternative or alternate may refer to: Arts, entertainment and media * Alternative (''Kamen Rider''), a character in the Japanese TV series ''Kamen Rider Ryuki'' * Alternative comics, or independent comics are an alternative to mainstream superh ...

or verticillate (in

whorls A whorl ( or ) is an individual circle, oval, volution or equivalent in a whorled pattern, which consists of a spiral or multiple concentric objects (including circles, ovals and arcs). In nature File:Photograph and axial plane floral diagram ...

). Leaves are usually

compound Compound may refer to: Architecture and built environments * Compound (enclosure), a cluster of buildings having a shared purpose, usually inside a fence or wall ** Compound (fortification), a version of the above fortified with defensive struc ...

bifoliate The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets) ...

trifoliate The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets ...

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...

, or

palmate The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets ...

, or rarely

simple Simple or SIMPLE may refer to: *Simplicity, the state or quality of being simple Arts and entertainment * ''Simple'' (album), by Andy Yorke, 2008, and its title track * "Simple" (Florida Georgia Line song), 2018 * "Simple", a song by John ...

Stipule In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole (botany), petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part ...

s are absent, but persistent; enlarged

axil A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, fl ...

lary

bud In botany, a bud is an undeveloped or Plant embryogenesis, embryonic Shoot (botany), shoot and normally occurs in the axil of a leaf or at the tip of a Plant stem, stem. Once formed, a bud may remain for some time in a dormancy, dormant conditi ...

scales Scale or scales may refer to: Mathematics * Scale (descriptive set theory), an object defined on a set of points * Scale (ratio), the ratio of a linear dimension of a model to the corresponding dimension of the original * Scale factor, a number ...

( pseudostipules) are often present.

Domatia A domatium (plural: domatia, from the Latin "domus", meaning home) is a tiny chamber that houses arthropods, produced by a plant. Ideally domatia differ from galls in that they are produced by the plant rather than being induced by their inhabi ...

occur in some genera. Dolichandrone falcata W2 IMG 9458

Flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...

s are solitary or in

inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...

s in a

raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...

or a

helicoid The helicoid, also known as helical surface, is a smooth Surface (differential geometry), surface embedded in three-dimensional space. It is the surface traced by an infinite line that is simultaneously being rotated and lifted along its Rotation ...

or dichasial cyme. Inflorescences bear persistent or

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

bract In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also lo ...

s or bractlets. The flowers are

hypogynous In the flowering plants, an ovary is a part of the female reproductive organ of the flower or gynoecium. Specifically, it is the part of the pistil which holds the ovule(s) and is located above or below or at the point of connection with the ba ...

zygomorphic Floral symmetry describes whether, and how, a flower, in particular its perianth, can be divided into two or more identical or mirror-image parts. Uncommonly, flowers may have no axis of symmetry at all, typically because their parts are spir ...

bisexual Bisexuality is romantic attraction, sexual attraction, or sexual behavior toward both males and females. It may also be defined as the attraction to more than one gender, to people of both the same and different gender, or the attraction t ...

, and usually conspicuous. The

calyx CALYX, Inc. is a non-profit publisher of art and literature by women founded in 1976 based in Corvallis, Oregon. CALYX publishes both '' CALYX, A Journal of Art and Literature by Women'' twice a year and CALYX Books, which publishes one to three ...

and corolla are distinct. The calyx is synsepalous, with five

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s. The corolla is

sympetalous Sympetally (fused petals) is a flower characteristic that historically was used to classify a grouping of plants termed Sympetalae, but this term has been abandoned in newer molecular based classifications, although the grouping has similarity to ...

, with five

petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol ...

s, often bilabiate. Corolla lobes are

imbricate Aestivation or estivation is the positional arrangement of the parts of a flower within a flower bud before it has opened. Aestivation is also sometimes referred to as praefoliation or prefoliation, but these terms may also mean vernation: the ar ...

in bud, or rarely valvate, and usually much shorter than the corolla tube.

Stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s are inserted on the corolla tube, alternating with corolla lobes. The four stamens are didynamous, members of each pair often

connivent Connation in plants is the developmental fusion of organs of the same type, for example, petals to one another to form a tubular corolla. This is in contrast to adnation, the fusion of dissimilar organs. Such organs are described as connate or a ...

, the

adaxial {{Short pages monitor Many species of Bignoniaceae have some use, either commercially or ethnobotanically, but the most important, by far, are those planted as

, especially the flowering trees. ''Jacaranda, Campsis, Pyrostegia, Tabebuia, Catalpa, Roseodendron, Handroanthus'' and ''Crescentia'' all have species of

horticultural Horticulture (from ) is the art and science of growing fruits, vegetables, flowers, trees, shrubs and ornamental plants. Horticulture is commonly associated with the more professional and technical aspects of plant cultivation on a smaller and mo ...

significance, at least in warm climates. Several others, including ''Tecoma, Podranea, Pandorea, Bignonia'' and ''Mansoa'' are frequently grown as ornamentals, at least in certain areas of the tropics. A great many species are known in cultivation, if only rarely. ''

Jacaranda mimosifolia ''Jacaranda mimosifolia'' is a sub-tropical tree native to south-central South America that has been widely planted elsewhere because of its attractive and long-lasting Violet (color), violet-colored flowers. It is also known as the jacaranda, b ...

'' is common as an avenue tree. The winged petiole and

leaf of ''

Crescentia alata ''Crescentia alata'', variously called Mexican calabash, jícaro, morro, morrito, or winged calabash, is a plant species in the family Bignoniaceae and in the genus ''Crescentia'', native to southern Mexico and Central America south to Costa Ric ...

'' resembles a

crucifixion Crucifixion is a method of capital punishment in which the condemned is tied or nailed to a large wooden cross, beam or stake and left to hang until eventual death. It was used as a punishment by the Achaemenid Empire, Persians, Ancient Carthag ...

cross, so is sometimes planted in the

Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...

as a

religious symbol A religious symbol is an iconic representation intended to represent a specific religion, or a specific concept within a given religion. Religious symbols have been used in the military in many countries, such as the United States military chap ...

. ''

Handroanthus ''Handroanthus'' is a genus of flowering plants in the Family (biology), family Bignoniaceae.David J. Mabberley. 2008. ''Mabberley's Plant-Book'' third edition (2008). Cambridge University Press: UK. It consists of 30 species of trees, known i ...

'' and the unrelated ''

Guaiacum ''Guaiacum'' (''OED'' 2nd edition, 1989.Entry "guaiacum"
in
Zygophyllaceae Zygophyllaceae is a family of flowering plants that contains the bean-caper and caltrop. The family includes around 285 species in 22 genera. Plants in the family Zygophyllaceae may be trees, shrubs, or herbs. They are often found in dry habit ...

) have the hardest, heaviest, and most durable wood of the American tropics. Important timber trees in ''Handroanthus'' include ''H. heptaphyllus'', ''H. serratifolius'', ''H. guayacan'', ''H. chrysanthus'', and ''H. billbergii''. ''

Tabebuia rosea ''Tabebuia rosea'', also called pink poui, and rosy trumpet tree is a neotropical tree that grows up to and can reach a diameter at breast height of up to . The Spanish name ''roble de sabana'', meaning "savannah oak", is widely used in Costa R ...

'' (including ''Tabebuia pentaphylla'') is harvested for

lumber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...

throughout the

New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...

tropics. ''

Tabebuia heterophylla ''Tabebuia heterophylla'' is a species of tree native to the Caribbean, and is also cultivated elsewhere. It is also known as roble blanco, pink manjack, pink trumpet tree, white cedar, and whitewood. Description ''Tabebuia heterophylla'' grows ...

'', and '' Tabebuia angustata'' are important sources of lumber for some of the

Caribbean islands Most of the Caribbean countries are islands in the Caribbean Sea, with only a few in inland lakes. The largest islands include Cuba, Hispaniola, Jamaica and Puerto Rico. Some of the smaller islands are referred to as a ''rock'' or ''reef.'' ''I ...

. Several species of ''Catalpa'' are also important

timber Lumber is wood that has been processed into uniform and useful sizes (dimensional lumber), including beams and planks or boards. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, window frames). ...

trees. '' Paratecoma'' was once the most important timber tree of the

Rio de Janeiro Rio de Janeiro, or simply Rio, is the capital of the Rio de Janeiro (state), state of Rio de Janeiro. It is the List of cities in Brazil by population, second-most-populous city in Brazil (after São Paulo) and the Largest cities in the America ...

area, but relentless exploitation has brought it to the verge of extinction. Several of the rare species of Bignoniaceae produce excellent wood, but are often not recognized by

lumberjack Lumberjack is a mostly North American term for workers in the logging industry who perform the initial harvesting and transport of trees. The term usually refers to loggers in the era before 1945 in the United States, when trees were felled us ...

s. Several uses of plants in Bignoniaceae are known locally. ''

Parmentiera aculeata ''Parmentiera aculeata'' (synonym ''Parmentiera edulis''), commonly known as cow okra, cuajilote, guajilote, huachilote, or pepino kat, is a species of flowering tree in the family Bignoniaceae. It is native to Mexico and Central America and is ...

'' is grown for its edible fruit in

Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...

and southern Mexico. The powdered seeds and sometimes the fruit pulp of ''

Crescentia cujete ''Crescentia cujete'', commonly known as the calabash tree, is a species of flowering plant a medium size tree in the trumpet vine family Bignoniaceae native to the Americas, and which is grown in Africa, Southeast Asia, Central America, So ...

'' and ''

'' are used in

Nicaragua Nicaragua, officially the Republic of Nicaragua, is the geographically largest Sovereign state, country in Central America, comprising . With a population of 7,142,529 as of 2024, it is the third-most populous country in Central America aft ...

to make a refresco called '' semilla de jicaro''.

Onion An onion (''Allium cepa'' , from Latin ), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the onion which was classifie ...

-scented species of ''Mansoa'' and

clove Cloves are the aromatic flower buds of a tree in the family Myrtaceae, ''Syzygium aromaticum'' (). They are native to the Maluku Islands, or Moluccas, in Indonesia, and are commonly used as a spice, flavoring, or Aroma compound, fragrance in fin ...

-scented species of ''Tynanthus'' are used as

condiment A condiment is a preparation that is added to food, typically after cooking, to enhance the Flavoring, flavour, to complement the dish or to impart a specific flavor. Such specific flavors generally add sweetness or pungency, or sharp or piquant ...

s. In northern

Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...

, shavings of the stems of '' Dolichandra quadrivalvis'' are added to bait which is left overnight near the burrows of

crab Crabs are decapod crustaceans of the infraorder Brachyura (meaning "short tailed" in Greek language, Greek), which typically have a very short projecting tail-like abdomen#Arthropoda, abdomen, usually hidden entirely under the Thorax (arthropo ...

s. The crabs are

paralyzed Paralysis (: paralyses; also known as plegia) is a loss of motor function in one or more muscles. Paralysis can also be accompanied by a loss of feeling (sensory loss) in the affected area if there is sensory damage. In the United States, r ...

for a few hours after eating the bait and are picked up by crabbers in the morning. The crabs recover before they reach market, and no harm from eating them has been reported. ''

Tanaecium nocturnum ''Tanaecium nocturnum'' (native name "kangara kane") is a species of flowering plant in the family Bignoniaceae, and is native to the Neotropics. Its white tubular flowers are long, and are pollinated by a hawkmoth with a tongue of similar lengt ...

'' is the source of a

hallucinogenic Hallucinogens, also known as psychedelics, entheogens, or historically as psychotomimetics, are a large and diverse class of psychoactive drugs that can produce altered states of consciousness characterized by major alterations in thought, moo ...

drug A drug is any chemical substance other than a nutrient or an essential dietary ingredient, which, when administered to a living organism, produces a biological effect. Consumption of drugs can be via insufflation (medicine), inhalation, drug i ...

.Christian Rätsch. 2005. ''The Encyclopedia of Psychoactive Plants'' (translated by John R. Baker). Park Street Press: Rochester VT, USA. . Its crushed leaves and stems are used to enervate

bees Bees are winged insects closely related to wasps and ants, known for their roles in pollination and, in the case of the best-known bee species, the western honey bee, for producing honey. Bees are a monophyletic lineage within the superfamil ...

while gathering

honey Honey is a sweet and viscous substance made by several species of bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of pl ...

. '' Fridericia chica'' is the source of a red

pigment A pigment is a powder used to add or alter color or change visual appearance. Pigments are completely or nearly solubility, insoluble and reactivity (chemistry), chemically unreactive in water or another medium; in contrast, dyes are colored sub ...

used in the

Amazon Basin The Amazon basin is the part of South America drained by the Amazon River and its tributary, tributaries. The Amazon drainage basin covers an area of about , or about 35.5 percent of the South American continent. It is located in the countries ...

for body paint and for

dye Juan de Guillebon, better known by his stage name DyE, is a French musician. He is known for the music video of the single "Fantasy Fantasy is a genre of speculative fiction that involves supernatural or Magic (supernatural), magical ele ...

in basketry. '' Cybistax antisyphilitica'' is the source of a blue dye commonly used in

Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...

. The bark of '' Sparattosperma leucantha'' is used in

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

to produce a brown dye for staining

cotton Cotton (), first recorded in ancient India, is a soft, fluffy staple fiber that grows in a boll, or protective case, around the seeds of the cotton plants of the genus '' Gossypium'' in the mallow family Malvaceae. The fiber is almost pure ...

thread. Medical claims are innumerable and usually spurious. Gentry describes an especially ludicrous example. Misidentification of plants, even by

botanist Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...

s, continues to be a big problem for ethnobotany, and it is especially severe for Bignoniaceae.

Voucher A voucher is a bond of the redeemable transaction type which is worth a certain money, monetary value and which may be spent only for specific reasons or on specific goods. Examples include house, housing, travel, and food vouchers. The term vou ...

specimen Specimen may refer to: Science and technology * Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount * Biological specimen or biospecimen, an organic specimen held by a biorepository f ...

s are often

sterile Sterile or sterility may refer to: *Asepsis, a state of being free from biological contaminants * Sterile (archaeology), a sediment deposit which contains no evidence of human activity *Sterilization (microbiology), any process that eliminates or ...

and fragmentary, making them nearly impossible to identify. False medical claims are often based on mistaken identification. The bark of several species of ''Handroanthus'' is sold in South American markets. Similar-looking bark is often fraudulently passed off as ''Handroanthus''. It is used in various ways to relieve certain

symptom Signs and symptoms are diagnostic indications of an illness, injury, or condition. Signs are objective and externally observable; symptoms are a person's reported subjective experiences. A sign for example may be a higher or lower temperature ...

s of certain

cancer Cancer is a group of diseases involving Cell growth#Disorders, abnormal cell growth with the potential to Invasion (cancer), invade or Metastasis, spread to other parts of the body. These contrast with benign tumors, which do not spread. Po ...

s. No evidence shows it prevents the disease or slows its progression, as is often claimed. '' Adenocalymma flavida'' has been used to relieve the aching of joints and muscles. A root

extract An extract (essence) is a substance made by extracting a part of a raw material, often by using a solvent such as ethanol, oil or water. Extracts may be sold as tinctures or absolutes or dried and powdered. The aromatic principles of ma ...

from '' Martinella'' is useful in the treatment of

conjunctivitis Conjunctivitis, also known as pink eye or Madras eye, is inflammation of the conjunctiva, the thin, clear layer that covers the white surface of the eye and the inner eyelid. It makes the eye appear pink or reddish. Pain, burning, scratchiness ...

and possibly other conditions of the eye.

References

Sources

* Alwyn H. Gentry. 1992. "Bignoniaceae: Part II (Tecomeae)". ''Flora Neotropica'' Monograph 25(2):1-150. (See ''External links'' below).