''Orthonama obstipata'', the gem, is a

moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...

of the family

Geometridae The geometer moths are moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyleti ...

. The species was first described by

Johan Christian Fabricius Johann Christian Fabricius (7 January 1745 – 3 March 1808) was a Danish zoologist, specialising in "Insecta", which at that time included all arthropods: insects, arachnids, crustaceans and others. He was a student of Carl Linnaeus, and is co ...

in 1794. It is a

cosmopolitan species In biogeography, a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats; most cosmopolitan species are known to be highly adaptable to a range of climatic and en ...

. In

continental Europe Continental Europe or mainland Europe is the contiguous mainland of Europe, excluding its surrounding islands. It can also be referred to ambiguously as the European continent, – which can conversely mean the whole of Europe – and, by som ...

though in the northeast, its range does not significantly extend beyond the

Baltic region The Baltic Sea Region, alternatively the Baltic Rim countries (or simply the Baltic Rim), and the Baltic Sea countries/states, refers to the general area surrounding the Baltic Sea, including parts of Northern, Central and Eastern Europe. Un ...

and it is absent from northern

Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...

. This well-flying species is prone to vagrancy and able to cross considerable distances of the open sea; it can thus be regularly found on the

British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...

(though mainly in the south) and even on

Iceland Iceland is a Nordic countries, Nordic island country between the Atlantic Ocean, North Atlantic and Arctic Oceans, on the Mid-Atlantic Ridge between North America and Europe. It is culturally and politically linked with Europe and is the regi ...

. Under its

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

s ''Nycterosea brunneipennis'' and ''Geometra fluviata'', the gem is the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...

''Nycterosea'' and ''Percnoptilota'', respectively. The latter is treated as a junior synonym of the former, but ''Nycterosea'', though usually included in ''Orthonama'' these days, may warrant recognition as an independent genus after all.

Description and ecology

The adult's

wingspan The wingspan (or just span) of a bird or an airplane is the distance from one wingtip to the opposite wingtip. For example, the Boeing 777–200 has a wingspan of , and a wandering albatross (''Diomedea exulans'') caught in 1965 had a wingsp ...

is ; in their core range (e.g.

Belgium Belgium, officially the Kingdom of Belgium, is a country in Northwestern Europe. Situated in a coastal lowland region known as the Low Countries, it is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeas ...

and the

Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...

) they can be seen between April and November, but in outlying regions, they may only be regularly encountered in late summer and early autumn, when vagrant individuals abound. This species is strongly

sexually dimorphic Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

: Males are light brown with a wavy pattern of whitish lines and a broad darker band running across the wings, forming

concentric In geometry, two or more objects are said to be ''concentric'' when they share the same center. Any pair of (possibly unalike) objects with well-defined centers can be concentric, including circles, spheres, regular polygons, regular polyh ...

semicircles when the moth is at rest. There is a small whitish-rimmed black spot within the darker band between the centre and the leading edge of each forewing. The females are slightly larger and much darker, almost uniformly blackish brown with an indistinct lighter pattern and a forewing spot like the males have. Adult caterpillars are greenish or brownish coloured and slightly tapered towards the head. The sides are yellowish, and from the middle body segments light, dark-lined diamond spots which are black cored stand out.

Similar species

Due to the very characteristic wing drawing, the female moths are unmistakable. In the males, there is a distant resemblance to grey-brown specimens of ''

Xanthorhoe designata ''Xanthorhoe designata'', the flame carpet, is a moth of the genus ''Xanthorhoe'' in the family Geometridae. The species was first described by Johann Siegfried Hufnagel in 1767. Distribution It is found in Europe, Asiatic Russia and Japan. D ...

''. In ''X. designata'' the midfield is, though, predominantly reddish-brown and bounded by a narrow black band.

Biology

The

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e feed on a wide range of low-growing

core eudicots The eudicots or eudicotyledons are flowering plants that have two seed leaves (cotyledons) upon germination. The term derives from ''dicotyledon'' (etymologically, ''eu'' = true; ''di'' = two; ''cotyledon'' = seed leaf). Historically, authors h ...

, but prefer

asterid Asterids are a large clade (monophyletic group) of flowering plants, composed of 17 orders and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked clades lamiids (8 orders) and ...

s. Host plants recorded from the

Central Europe Central Europe is a geographical region of Europe between Eastern Europe, Eastern, Southern Europe, Southern, Western Europe, Western and Northern Europe, Northern Europe. Central Europe is known for its cultural diversity; however, countries in ...

an part of its range include:

Euasterids I Asterids are a large clade (monophyletic group) of flowering plants, composed of 17 orders and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked clades lamiids (8 orders) and ...

Gentianales Gentianales is an order of flowering plant, included within the asterid clade of eudicots. It comprises more than 20,000 species in about 1,200 genera in 5 families. More than 80% of the species in this order belong to the family Rubiaceae. Man ...

Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with Petiole ( ...

** ''

Galium ''Galium'' is a large genus of annual and perennial herbaceous plants in the family Rubiaceae, occurring in the temperate zones of both the Northern and Southern Hemispheres. Some species are informally known as bedstraw. Over 600 species of ' ...

'' (bedstraws) *

Solanales The Solanales are an order of flowering plants, included in the asterid group of the eudicots. Well-known members of Solanales include potatoes, eggplants, tomatoes, chili peppers, tobacco, petunias, nightshades, morning glory, and sweet potato. ...

Convolvulaceae Convolvulaceae (), commonly called the bindweed, bindweeds or morning glory, morning glories, is a Family (biology), family of about 60 genera and more than 1,650 species. These species are primarily herbaceous vines, but also include trees, sh ...

** ''

Convolvulus ''Convolvulus'' is a genus of about 200 to 250''Convolvulus''.
Flora of China.

'' (true bindweeds)

Euasterids II Asterids are a large clade (monophyletic group) of flowering plants, composed of 17 orders and more than 80,000 species, about a third of the total flowering plant species. The asterids are divided into the unranked clades lamiids (8 orders) an ...

Asterales Asterales ( ) is an Order (biology), order of dicotyledonous flowering plants that includes the large Family (biology), family Asteraceae (or Compositae) known for composite flowers made of Floret#floret, florets, and ten families related to th ...

Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...

** ''

Anthemis ''Anthemis'' is a genus of aromatic flowering plants in the family (biology), family Asteraceae, closely related to ''Chamaemelum'', and like that genus, known by the common name chamomile; some species are also called dog-fennel or mayweed. ''An ...

'' (dog-fennels) ** ''

Chrysanthemum Chrysanthemums ( ), sometimes called mums or chrysanths, are flowering plants in the Asteraceae family. They are native to East Asia and northeastern Europe. Most species originate from East Asia, and the center of diversity is in China. Co ...

'' (chrysanthemums) ** ''

Eupatorium ''Eupatorium'' is a genus of flowering plants in the family Asteraceae, containing from 36 to 60 species depending on the classification system. Most are Herbaceous plant, herbaceous perennial plant, perennials growing to tall. A few are shrubs ...

'' (bonesets) ** ''

Senecio ''Senecio'' is a genus of flowering plants in the daisy family (Asteraceae) that includes ragworts and groundsels. Variously circumscribed taxonomically, the genus ''Senecio'' is one of the largest genera of flowering plants. Description Mo ...

'' (ragworts and groundsels) – e.g. common groundsel (''S. vulgaris'') in the

Eurosids II The rosids are members of a large clade (monophyletic group) of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classificatio ...

Brassicales The Brassicales (or Cruciales) are an order (biology), order of flowering plants, belonging to the malvid group of eudicotyledons under the APG IV system. Well-known members of Brassicales include cabbage, cauliflower, Brussels sprout, broccoli, ...

Brassicaceae Brassicaceae () or (the older but equally valid) Cruciferae () is a medium-sized and economically important Family (biology), family of flowering plants commonly known as the mustards, the crucifers, or the cabbage family. Most are herbaceous pla ...

** ''

Alyssum ''Alyssum'' is a genus of over a hundred species of flowering plants in the family Brassicaceae, native to Europe, Asia, and northern Africa, with the highest species diversity in the Mediterranean region. The genus comprises annual plant, annu ...

'' (alyssums) ** '' Nasturtium'' (watercresses) Basal core eudicots *

Caryophyllales Caryophyllales ( ) is a diverse and heterogeneous order of flowering plants with well-known members including cacti, carnations, beets, quinoa, spinach, amaranths, pigfaces and ice plants, oraches and saltbushes, goosefoots, sundews, Venu ...

Polygonaceae The Polygonaceae are a Family (biology), family of flowering plants known Common name, informally as the knotweed family or smartweed—buckwheat family in the United States. The Botanical name, name is Basionym, based on the genus ''Polygonum'', ...

** ''

Polygonum ''Polygonum'' is a genus of about 130 species of flowering plants in the buckwheat and knotweed family Polygonaceae. Common names include knotweed and knotgrass (though the common names may refer more broadly to plants from Polygonaceae). In the ...

'' (knotweed) ** ''

Rumex The docks and sorrels, genus ''Rumex'', are a genus of about 200 species of annual, biennial, and perennial herbs in the buckwheat family, Polygonaceae. Members of this genus are very common perennial herbs with a native almost worldwide distri ...

'' (docks and sorrels)

Synonyms

The widespread, strongly

and somewhat

phenotypical In genetics, the phenotype () is the set of observable characteristics or phenotypic trait, traits of an organism. The term covers the organism's morphology (biology), morphology (physical form and structure), its Developmental biology, develo ...

ly variable gem has been described anew time and again by various authors, even as late as the early 20th century, but all these supposedly distinct

taxa In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...

are nowadays considered to refer to a single species.

Junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

s of the gem include:Pitkin & Jenkins (2004ab), and see references in Savela (2002)

Footnotes

References

* (1942)
"Seltsame Geometridenfunde"
eculiar records of geometer moths ''Zeitschrift des Wiener Entomologen-Vereins''. 27: 109.
"70.047 BF1720 The Gem ''Nycterosea obstipata'' (Fabricius, 1794)"
Retrieved May 15, 2019. * * *

External links

''Nycterosea obstipata'' on ''Fauna Europaea''''Lepiforum e.V.''''Nycterosea obstipata'' on ''De Vlinderstichting''
{{DEFAULTSORT:Orthonama Obstipata Xanthorhoini Moths of Europe Moths of Cape Verde Moths of Africa Moths of Iceland Moths of Japan Moths described in 1794 Taxa named by Johan Christian Fabricius