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The nasal cavity is a large, air-filled space above and behind the
nose A nose is a protuberance in vertebrates that houses the nostrils, or nares, which receive and expel air for respiration alongside the mouth. Behind the nose are the olfactory mucosa and the sinuses. Behind the nasal cavity, air next pass ...
in the middle of the face. The nasal septum divides the cavity into two cavities, also known as fossae. Each cavity is the continuation of one of the two
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called t ...
s. The nasal cavity is the uppermost part of the
respiratory system The respiratory system (also respiratory apparatus, ventilatory system) is a biological system consisting of specific organs and structures used for gas exchange in animals and plants. The anatomy and physiology that make this happen varies grea ...
and provides the nasal passage for inhaled air from the
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called t ...
s to the nasopharynx and rest of the respiratory tract. The paranasal sinuses surround and drain into the nasal cavity.


Structure

The term "nasal cavity" can refer to each of the two cavities of the nose, or to the two sides combined. The lateral wall of each nasal cavity mainly consists of the
maxilla The maxilla (plural: ''maxillae'' ) in vertebrates is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The ...
. However, there is a deficiency that is compensated for by the perpendicular plate of the palatine bone, the medial pterygoid plate, the labyrinth of ethmoid and the inferior concha. The paranasal sinuses are connected to the nasal cavity through small orifices called
ostia Ostia may refer to: Places *Ostia (Rome), a municipio (also called ''Ostia Lido'' or ''Lido di Ostia'') of Rome *Ostia Antica, a township and port of ancient Rome *Ostia Antica (district), a district of the commune of Rome Arts and entertainment ...
. Most of these ostia communicate with the nose through the lateral nasal wall, via a semi-lunar depression in it known as the semilunar hiatus. The hiatus is bound laterally by a projection known as the uncinate process. This region is called the ostiomeatal complex. The roof of each nasal cavity is formed in its upper third to one half by the nasal bone and more inferiorly by the junctions of the upper lateral cartilage and nasal septum. Connective tissue and skin cover the bony and cartilaginous components of the nasal dorsum. The floor of the nasal cavities, which also form the roof of the mouth, is made up by the bones of the hard palate: the horizontal plate of the palatine bone posteriorly and the palatine process of the maxilla anteriorly. The most anterior part of the nasal cavity is the nasal vestibule. The vestibule is enclosed by the cartilages of the nose and lined by the same epithelium of the
skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other cuticle, animal coverings, such as the arthropod exoskeleton, have diffe ...
(stratified squamous, keratinized). Within the vestibule this changes into the typical respiratory epithelium that lines the rest of the nasal cavity and respiratory tract. Inside the
nostril A nostril (or naris , plural ''nares'' ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called t ...
s of the vestibule are the nasal hair, which filter dust and other matter that are breathed in. The back of the cavity blends, via the choanae, into the nasopharynx. The nasal cavity is divided in two by the vertical nasal septum. On the side of each nasal cavity are three horizontal outgrowths called nasal conchae (singular "concha") or turbinates. These turbinates disrupt the airflow, directing air toward the olfactory epithelium on the surface of the turbinates and the septum. The vomeronasal organ is located at the back of the
septum In biology, a septum (Latin for ''something that encloses''; plural septa) is a wall, dividing a cavity or structure into smaller ones. A cavity or structure divided in this way may be referred to as septate. Examples Human anatomy * Interat ...
and has a role in pheromone detection.


Segments

The nasal cavity is divided into two segments: the respiratory segment and the olfactory segment. * The respiratory segment comprises most of each nasal cavity, and is lined with ciliated pseudostratified columnar epithelium (also called respiratory epithelium). The conchae, or turbinates, are located in this region. The turbinates have a very vascularized lamina propria (erectile tissue) allowing the venous plexuses of their mucosa to engorge with blood, restricting airflow and causing air to be directed to the other side of the nose, which acts in concert by shunting blood out of its turbinates. This
cycle Cycle, cycles, or cyclic may refer to: Anthropology and social sciences * Cyclic history, a theory of history * Cyclical theory, a theory of American political history associated with Arthur Schlesinger, Sr. * Social cycle, various cycles in soc ...
occurs approximately every two and a half hours. * The olfactory segment is lined with a specialized type of pseudostratified columnar epithelium, known as olfactory epithelium, which contains receptors for the sense of the smell. This segment is located in and beneath the mucosa of the roof of each nasal cavity and the medial side of each middle turbinate. Histological sections appear yellowish-brown due to the presence of lipofuscin pigments. Olfactory mucosal cell types include bipolar neurons, supporting (sustentacular) cells, basal cells, and Bowman's glands. The axons of the bipolar neurons form the olfactory nerve (cranial nerve I) which enters the brain through the cribriform plate. Bowman's glands are serous glands in the lamina propria, whose secretions trap and dissolve odoriferous substances.


Blood supply

There is a rich blood supply to the nasal cavity. In some animals, such as dogs, the capillary beds flowing through the nasal cavity help cool the blood flow to the
brain The brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It consists of nervous tissue and is typically located in the head ( cephalization), usually near organs for special ...
. Blood supply comes from branches of both the internal and external carotid artery, including branches of the
facial artery The facial artery (external maxillary artery in older texts) is a branch of the external carotid artery that supplies structures of the superficial face. Structure The facial artery arises in the carotid triangle from the external carotid arter ...
and maxillary artery. The named arteries of the nose are: * Sphenopalatine artery and greater palatine artery, branches of the maxillary artery. * Anterior ethmoidal artery and posterior ethmoidal artery, branches of the ophthalmic artery *Septal branches of the superior labial artery, a branch of the
facial artery The facial artery (external maxillary artery in older texts) is a branch of the external carotid artery that supplies structures of the superficial face. Structure The facial artery arises in the carotid triangle from the external carotid arter ...
, which supplies the vestibule of the nasal cavity.


Nerve supply

Innervation A nerve is an enclosed, cable-like bundle of nerve fibers (called axons) in the peripheral nervous system. A nerve transmits electrical impulses. It is the basic unit of the peripheral nervous system. A nerve provides a common pathway for the ...
of the nasal cavity responsible for the sense of smell is via the olfactory nerve, which sends microscopic fibers from the olfactory bulb through the cribriform plate to reach the top of the nasal cavity. General sensory innervation is by branches of the
trigeminal nerve In neuroanatomy, the trigeminal nerve (literal translation, lit. ''triplet'' nerve), also known as the fifth cranial nerve, cranial nerve V, or simply CN V, is a cranial nerve responsible for Sense, sensation in the face and motor functions ...
(V1 & V2): * Nasociliary nerve (V1) * Anterior Ethmoidal nerve from the nasociliary nerve (V1) *Posterior nasal branches of Maxillary nerve (V2) The nasal cavity is innervated by autonomic fibers. Sympathetic innervation to the blood vessels of the mucosa causes them to
constrict Constriction is a method used by various snake species to kill or subdue their prey. Although some species of venomous and mildly venomous snakes do use constriction to subdue their prey, most snakes which use constriction lack venom. The snake i ...
, while the control of secretion by the mucous glands is carried on postganglionic
parasympathetic The parasympathetic nervous system (PSNS) is one of the three divisions of the autonomic nervous system, the others being the sympathetic nervous system and the enteric nervous system. The enteric nervous system is sometimes considered part o ...
nerve fibers originating from the
facial nerve The facial nerve, also known as the seventh cranial nerve, cranial nerve VII, or simply CN VII, is a cranial nerve that emerges from the pons of the brainstem, controls the muscles of facial expression, and functions in the conveyance of ta ...
.


Function

The two nasal cavities condition the air to be received by the other areas of the respiratory tract. Owing to the large surface area provided by the nasal conchae (also known as turbinates), the air passing through the nasal cavity is warmed or cooled to within 1 degree of
body temperature Thermoregulation is the ability of an organism to keep its body temperature within certain boundaries, even when the surrounding temperature is very different. A thermoconforming organism, by contrast, simply adopts the surrounding temperatur ...
. In addition, the air is humidified, and dust and other particulate matter is removed by nasal hair in the nostrils. The entire mucosa of the nasal cavity is covered by a blanket of mucus, which lies superficial to the microscopic cilia and also filters inspired air. The
cilia The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projecti ...
of the respiratory epithelium move the secreted mucus and particulate matter posteriorly towards the
pharynx The pharynx (plural: pharynges) is the part of the throat behind the mouth and nasal cavity, and above the oesophagus and trachea (the tubes going down to the stomach and the lungs). It is found in vertebrates and invertebrates, though its ...
where it passes into the
esophagus The esophagus (American English) or oesophagus (British English; both ), non-technically known also as the food pipe or gullet, is an organ in vertebrates through which food passes, aided by peristaltic contractions, from the pharynx to the ...
and is digested in the stomach. The nasal cavity also houses the sense of smell and contributes greatly to taste sensation through its posterior communication with the mouth via the choanae.


Clinical significance

Diseases of the nasal cavity include
viral Viral means "relating to viruses" (small infectious agents). Viral may also refer to: Viral behavior, or virality Memetic behavior likened that of a virus, for example: * Viral marketing, the use of existing social networks to spread a marke ...
,
bacteria Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
l and fungal infections, nasal cavity tumors, both benign and much more often malignant, as well as inflammations of the nasal mucosa. Many problems can affect the nose, including: * Deviated septum - a shifting of the wall that divides the nasal cavity into halves * Nasal polyps - soft growths that develop on the lining of the nose or sinuses * Nosebleeds * Rhinitis - inflammation of the nose and sinuses sometimes caused by allergies. The main symptom is a runny nose. * Nasal fractures, also known as a broken nose * Common cold * Sinonasal tumors


See also

* Nasal irrigation * Danger triangle of the face


References


External links

* * Gross anatomy dissection of the nasal cavity, vide

an

{{DEFAULTSORT:Nasal Cavity Nose Olfactory system Respiratory system Sensory organs