''Moschops'' (Greek for "calf face") is an
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
therapsids that lived in the
Guadalupian
The Guadalupian is the second and middle Series (stratigraphy), series/Epoch (geology), epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico an ...
epoch
In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured.
The moment of epoch is usually decided b ...
, around 265–260 million years ago. They were heavily built plant eaters, and they may have lived partly in water, as
hippopotamus
The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic mammal native to sub-Sahar ...
es do. They had short, thick heads and might have competed by head-butting each other. Their elbow joints allowed them to walk with a more
mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
-like gait rather than crawling. Their remains were found in the
Karoo
The Karoo ( ; from the Afrikaans borrowing of the South Khoekhoe Khoemana (also known as !Orakobab or Korana) word is a semidesert natural region of South Africa. No exact definition of what constitutes the Karoo is available, so its extent is ...
region of
South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, belonging to the
''Tapinocephalus'' Assemblage Zone. Therapsids, such as ''Moschops'', are
synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...
s, the dominant
land animal
Terrestrial animals are animals that live predominantly or entirely on land (e.g. cats, chickens, ants, most spiders), as compared with aquatic animals, which live predominantly or entirely in the water (e.g. fish, lobsters, octopuses), and semi ...
s in the
Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...
period, which ended 252 million years ago.
Description

''Moschops'' were heavy set
dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. ...
n
synapsid
Synapsida is a diverse group of tetrapod vertebrates that includes all mammals and their extinct relatives. It is one of the two major clades of the group Amniota, the other being the more diverse group Sauropsida (which includes all extant rept ...
s, measuring in length,
and weighing on average and in maximum body mass.
They had small heads with broad
orbits
In celestial mechanics, an orbit (also known as orbital revolution) is the curved trajectory of an physical body, object such as the trajectory of a planet around a star, or of a natural satellite around a planet, or of an satellite, artificia ...
and heavily built short necks. Like other members of
Tapinocephalidae, the skull had a tiny opening for the
pineal organ.
The Age of Reptiles
/ref> The occiput
The occipital bone () is a cranial dermal bone and the main bone of the occiput (back and lower part of the skull). It is trapezoidal in shape and curved on itself like a shallow dish. The occipital bone lies over the occipital lobes of the ...
was broad and deep, but the skull was more narrow in the dorsal border. Furthermore, the pterygoid arches and the angular region of the jaw with heavily built jaw muscles. Due to that and the possession of long-crowned, stout teeth, it is believed that ''Moschops'' was a herbivore feeding on nutrient-poor and tough vegetation, like cycad
Cycads are seed plants that typically have a stout and woody (ligneous) trunk (botany), trunk with a crown (botany), crown of large, hard, stiff, evergreen and (usually) pinnate leaves. The species are dioecious, that is, individual plants o ...
stems. Due to the presumably nutrient-poor food, it is likely they had to feed for long periods of time. The anatomy of the taxa allowed them to open the elbow joints more widely, enabling them to move in a more mammal-like posture than some other animals at the time. This helped to carry their massive bodies more easily while feeding, as well as allowing them short bursts of speed. It has also been proposed that ''Moschops'' were possibly sub-aquatic. ''Moschops'' had rather thick skulls, prompting speculation that individuals could have competed with one another by head-butting. A 2017 published study would later confirm this by synchrotron
A synchrotron is a particular type of cyclic particle accelerator, descended from the cyclotron, in which the accelerating particle beam travels around a fixed closed-loop path. The strength of the magnetic field which bends the particle beam i ...
scanning a ''Moschops capensis'' skull, which revealed numerous anatomical adaptations to the central nervous system for combative behaviour. They were likely preyed upon by titanosuchids and larger therocephalian species.
Earliest finds
''Moschops'' material was first discovered in the Ecca Group (part of the Karoo Supergroup
The Karoo Supergroup is the most widespread stratigraphic unit in Africa south of the Kalahari Desert. The supergroup consists of a sequence of units, mostly of nonmarine origin, deposited between the Late Carboniferous and Early Jurassic, a per ...
) of South Africa
South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
by Robert Broom
Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...
. As the geological horizon was dubious, it was referred to have originated from the Ecca Group on the basis of ''Pareiasaurus
''Pareiasaurus'' (from , "cheek" and , "lizard") is an extinct genus of Pareiasauromorpha, pareiasauromorph reptile from the Permian period. It was a typical member of its family (biology), family, the pareiasaurids, which take their name from th ...
'' remains in near proximity. The discovered material includes a holotype
A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
(AMNH 5550) and seven topotypes (AMNH 5551-5557). The degree of pachyostosis
Pachyostosis is a non-pathological condition in vertebrate animals in which the bones experience a thickening, generally caused by extra layers of lamellar bone. It often occurs together with bone densification ( osteosclerosis), reducing inner c ...
varies within the skulls of the specimens, and Broom believed this to have been linked to variations in gender and age. In 1910, the material was sent to the American Museum of Natural History
The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...
in New York City
New York, often called New York City (NYC), is the most populous city in the United States, located at the southern tip of New York State on one of the world's largest natural harbors. The city comprises five boroughs, each coextensive w ...
and described in 1911.
Classification
''Moschops'' is characterized by a strongly pachyostosed skull with a broad intertemporal region and greatly reduced temporal fossa
The temporal fossa is a fossa (shallow depression) on the side of the skull bounded by the temporal lines above, and the zygomatic arch below. Its floor is formed by the outer surfaces of four bones of the skull. The fossa is filled by the te ...
e. Two species are known from the fossil record, ''M. capensis'' and ''M. koupensis''. Two other species were assigned (''M. whaitsi'' and ''M. oweni''), but their validity is considered possibly dubious. Genera regarded as synonyms are ''Moschoides'', ''Agnosaurus'', ''Moschognathus'' and ''Pnigalion''. ''Delphinognathus conocephalus'' could represent juvenile ''Moschops'', thus possibly synonymous. ''Delphinognathus'' is only known from a single, moderately pachyostosed skull. It has a conical boss on the parietal surrounding the pineal foramen.
See also
*
References
External links
''Moschops''
pictures and a brief overview
at Paleos.com
{{Taxonbar, from=Q130737
Tapinocephalia
Prehistoric therapsid genera
Guadalupian synapsids of Africa
Permian South Africa
Fossils of South Africa
Fossil taxa described in 1911
Taxa named by Robert Broom