Morphology (biology)
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Morphology (from Ancient Greek μορφή (morphḗ) "form", and λόγος (lógos) "word, study, research") is the study of the form and structure of
organism An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...
s and their specific structural features. This includes aspects of the outward appearance (shape, structure, color, pattern, size), as well as the form and structure of internal parts like bones and organs, i.e., anatomy. This is in contrast to physiology, which deals primarily with function. Morphology is a branch of life science dealing with the study of the overall structure of an
organism An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have be ...
or
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
and its component parts.


History

The etymology of the word "morphology" is from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
(), meaning "form", and (), meaning "word, study, research". While the concept of form in biology, opposed to function, dates back to Aristotle (see
Aristotle's biology Aristotle's biology is the theory of biology, grounded in systematic observation and collection of data, mainly zoology, zoological, embodied in Aristotle's books on the science. Many of his observations were made during his stay on the island ...
), the field of morphology was developed by Johann Wolfgang von Goethe (1790) and independently by the German anatomist and physiologist Karl Friedrich Burdach (1800). Among other important theorists of morphology are Lorenz Oken,
Georges Cuvier Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
, Étienne Geoffroy Saint-Hilaire, Richard Owen, Carl Gegenbaur and Ernst Haeckel. In 1830, Cuvier and Saint-Hilaire engaged in a famous debate, which is said to exemplify the two major deviations in biological thinking at the time – whether animal structure was due to function or evolution.


Divisions of morphology

* Comparative morphology is an analysis of the patterns of the locus of structures within the body plan of an organism, and forms the basis of taxonomical categorization. * Functional morphology is the study of the relationship between the structure and function of morphological features. * Experimental morphology is the study of the effects of external factors upon the morphology of organisms under experimental conditions, such as the effect of genetic mutation. * Anatomy is a "branch of morphology that deals with the structure of organisms". * Molecular morphology is a rarely used term, usually referring to the superstructure of polymers such as fiber formation or to larger composite assemblies. The term is commonly not applied to the spatial structure of individual
molecules A molecule is a group of two or more atoms that are held together by attractive forces known as chemical bonds; depending on context, the term may or may not include ions that satisfy this criterion. In quantum physics, organic chemistry ...
. * Gross morphology refers to the collective structures of an organism as a whole as a general description of the form and structure of an organism, taking into account all of its structures without specifying an individual structure.


Morphology and classification

Most taxa differ morphologically from other taxa. Typically, closely related taxa differ much less than more distantly related ones, but there are exceptions to this. Cryptic species are
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
which look very similar, or perhaps even outwardly identical, but are reproductively isolated. Conversely, sometimes unrelated taxa acquire a similar appearance as a result of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
or even
mimicry In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
. In addition, there can be morphological differences within a species, such as in '' Apoica flavissima'' where queens are significantly smaller than workers. A further problem with relying on morphological data is that what may appear morphologically to be two distinct species may in fact be shown by DNA analysis to be a single species. The significance of these differences can be examined through the use of allometric engineering in which one or both species are manipulated to phenocopy the other species. A step relevant to the evaluation of morphology between traits/features within species, includes an assessment of the terms: homology an
homoplasy
Homology between features indicates that those features have been derived from a common ancestor. Alternatively, homoplasy between features describes those that can resemble each other, but derive independently via parallel or
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
.


3D cell morphology: classification

The invention and development of microscopy enabled the observation of 3-D cell morphology with both high spatial and temporal resolution. The dynamic processes of this cell morphology which are controlled by a
complex system A complex system is a system composed of many components that may interact with one another. Examples of complex systems are Earth's global climate, organisms, the human brain, infrastructure such as power grid, transportation or communication sy ...
play an important role in varied important biological processes, such as immune and invasive responses.


See also

* Comparative anatomy * Computational anatomy * Insect morphology * Morphometrics * Neuromorphology * Phenetics * Phenotype * Phenotypic plasticity * Plant morphology


References


External links

* {{Authority control Branches of biology Comparative anatomy