''Montanoceratops'' is an extinct

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of small

ceratopsian Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period, although ancestral forms lived earlier, in the Jurassi ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

that lived approximately 70

million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...

during the latter part of the

Cretaceous The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...

Period in what is now Montana and Alberta. ''Montanoceratops'' was a small sized, moderately-built, ground-dwelling,

quadrupedal Quadrupedalism is a form of locomotion where four limbs are used to bear weight and move around. An animal or machine that usually maintains a four-legged posture and moves using all four limbs is said to be a quadruped (from Latin ''quattuo ...

herbivore A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...

, that could grow up to an estimated in length and in body mass.

Description

''Montanoceratops'' was a typical primitive ceratopsian in many respects, distinguished from the later species by the presence of claws, rather than hooves, and by having teeth in its upper jaw, rather than a toothless

beak The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for fo ...

. It was once thought to have a horn on its nose but that was a misplaced cheek horn.Holtz, Thomas R. Jr. (2008) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages'
Supplementary Information
/ref> According to Brown and Schlaikjer (1935), ''Montanoceratops'' can be distinguished based on the following characteristics: * the nasal bone is proportionally large, deep, heavy and with a very well developed horncore * the

dentary In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bone ...

is long with a straight ventral margin *

Discovery and species

The first

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...

remains of what we now know as ''Montanoceratops'' were discovered on the Blackfeet Indian Reservation, west of Buffalo Lake,

Montana Montana () is a U.S. state, state in the Mountain states, Mountain West List of regions of the United States#Census Bureau-designated regions and divisions, division of the Western United States. It is bordered by Idaho to the west, North ...

in the

St. Mary River Formation The St. Mary River Formation is a geologic formation of Late Cretaceous (71.9-67 Ma) age of the Western Canada Sedimentary Basin in southwestern Alberta and northwesternmost Montana. It was first described from outcrops along the St. Mary River ...

. It was collected in 1916 by

Barnum Brown Barnum Brown (February 12, 1873 – February 5, 1963), commonly referred to as Mr. Bones, was an American paleontologist. Named after the circus showman P. T. Barnum, he discovered the first documented remains of ''Tyrannosaurus'' during a career ...

and Peter C. Kaisen of the American Museum of Natural History from terrestrial sediments that were deposited during the

Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...

stages Stage or stages may refer to: Acting * Stage (theatre), a space for the performance of theatrical productions * Theatre, a branch of the performing arts, often referred to as "the stage" * ''The Stage'', a weekly British theatre newspaper * St ...

of the

period, approximately 70 million years ago. After having mounted a reconstruction in 1935, Brown and his assistant Erich Maren Schlaikjer named it in 1942 as a new species of ''

Leptoceratops ''Leptoceratops'' (meaning 'Thin-horned face' and derived from Greek ''lepto-/λεπτο-'' meaning 'small', 'insignificant', 'slender', 'meagre' or 'lean', ''kerat-/κερατ-'' meaning 'horn' and ''-ops/ωψ'' meaning face), is a genus of lep ...

'': ''Leptoceratops cerorhynchus''; this is the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

of ''Montanoceratops''. The genus name ''Montanoceratops'', means "Montana horned face", and is derived from the state of Montana, which is the site of the discovery of the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

specimen, and the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

words "keras" (κερας) meaning "horn", and "ops" (ωψ) meaning "face". The

type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...

and only valid

species In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...

known today is ''Montanoceratops cerorhynchus''. The original type material discovered by Barnum Brown, designated specimen AMNH 5464, included an incomplete skull and mandible (with most of the skull absent), a complete series of eleven cervical, twelve dorsal and eight sacral vertebrae, thirteen complete caudal vertebrae and the centra of two others, several ribs, a complete pelvic girdle except for the right pubis and the distal part of the right ischium, both femora (346mm), the left tibia (355mm), left fibula and left astragalus, the second phalanx of digit three, and the ungual phalanges of the first, third and fourth digits of the left pes (foot). This specimen is housed in the collection of the American Museum of Natural History in

New York City New York, often called New York City or NYC, is the most populous city in the United States. With a 2020 population of 8,804,190 distributed over , New York City is also the most densely populated major city in the U ...

, USA. In 1986,

David B. Weishampel Professor David Bruce Weishampel (born November 16, 1952) is an American palaeontologist in the Center for Functional Anatomy and Evolution at Johns Hopkins University School of Medicine. Weishampel received his Ph.D. in Geology from the Univers ...

discovered more material referable to ''Montanoceratops'' in the Little Rocky Coulee locality of the St. Mary River Formation, in

Glacier County Glacier County is located in the U.S. state of Montana. As of the 2020 census, the population was 13,778. The county is located in northwestern Montana between the Great Plains and the Rocky Mountains, known to the Blackfeet as the "Backbone ...

, Montana. The material, which was found in terrestrial sediments, was also considered Maastrichtian in age. A description of this material was published in a paper by Brenda Chinnery and Weishampel in 1998. In 2001, Peter Makovicky examined and described specimen AMNH 5244, an incomplete but well-preserved

braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...

that he assigned to ''Montanoceratops''. This specimen was collected by Barnum Brown in 1910, from terrestrial sediments on the east bank of the Red Deer River near Tolman Ferry at the

Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of the ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...

.P. J. Makovicky. 2001. A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta. Mesozoic Vertebrate Life, D. H. Tanke and K. Carpenter (eds.), Indiana University Press, Bloomington 243-262 This specimen is also housed in the collection of the American Museum of Natural History.

Classification

In 1942, Brown assigned the type material AMNH 5464 to the genus ''

'' which he concluded was a member of the

taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...

"protoceratopsidae". In 1951,

Charles Mortram Sternberg Charles Mortram Sternberg (1885–1981) was an American-Canadian fossil collector and paleontologist, son of Charles Hazelius Sternberg. Late in his career, he collected and described '' Pachyrhinosaurus'', '' Brachylophosaurus'', '' Parksosau ...

examined more material belonging to ''Leptoceratops'' which showed that AMNH 5464 was a distinct genus, and hence he reassigned it to a new genus ''Montanoceratops''. In 1996, Chinnery and Weishampel conducted a phylogenetic analysis of basal neoceratopsians which showed that the "protoceratopsidae" was a

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...

group (i.e. not a valid taxon) and designated ''Montanoceratops'' as the most advanced basal neoceratopsian. In 2001, Mackovicky, concluded that ''Montanoceratops'' belonged to the taxon

Leptoceratopsidae Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, b ...

, and defined this group as consisting of ''

gracilis'' and all species closer to ''Leptoceratops'' than to ''Triceratops horridus''. A study by Ryan et al. (2012) confirmed this assignment. The following

cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...

is based on a 2015 analysis by Yiming He, Peter J. Makovicky, Kebai Wang, Shuqing Chen, Corwin Sullivan, Fenglu Han, Xing XuMichael J. Ryan, David C. Evans, Philip J. Currie, Caleb M. Brown and Don Brinkman.

Paleobiology

Another unusual feature was the presence of tall spines on the bones of the tail. Although these would not have been visible during life, they would have made the tail unusually deep in cross-section. Since the tail was also highly flexible, it is possible that it was used in intra-species signalling, and that the deep shape made it more visible. ''Montanoceratops'', like all Ceratopsians, was a

. It would have used its sharp Ceratopsian beak to bite off the leaves or needles.

Paleopathology

In specimen AMNH 5464, the neural spines of several of the

caudal vertebrae The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic ...

show an

ankylosis Ankylosis is a stiffness of a joint due to abnormal adhesion and rigidity of the bones of the joint, which may be the result of injury or disease. The rigidity may be complete or partial and may be due to inflammation of the tendinous or muscular ...

that is a result of a severe injury sustained by this individual during its lifetime. Evidence of this injury is also present in the outer surface of the left

ischium The ischium () form ...

, where there is a large, irregularly shaped growth of bone that has formed across a healed fracture.

Paleoecology

Habitat

The St. Mary River Formation has not undergone a definitive radiometric dating, however, the available stratigraphic correlation has shown that this formation was laid down between 74 and 66 million years ago during the final regression of the mid-continental Bearpaw Seaway. It ranges from as far south as Glacier County,

to as far north as the Little Bow River in Alberta. The St. Mary River Formation is part of the

Western Canadian Sedimentary Basin The Western Canadian Sedimentary Basin (WCSB) underlies of Western Canada including southwestern Manitoba, southern Saskatchewan, Alberta, northeastern British Columbia and the southwest corner of the Northwest Territories. This vast sedimentary ...

in southwestern Alberta, which extends from the

Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico in ...

in the west to the

Canadian Shield The Canadian Shield (french: Bouclier canadien ), also called the Laurentian Plateau, is a geologic shield, a large area of exposed Precambrian igneous and high-grade metamorphic rocks. It forms the North American Craton (or Laurentia), the anc ...

in the east. It is laterally equivalent to the

. The region where dinosaurs lived was bounded by mountains to the west, and included ancient channels, small freshwater ponds, streams, and

floodplain A floodplain or flood plain or bottomlands is an area of land adjacent to a river which stretches from the banks of its channel to the base of the enclosing valley walls, and which experiences flooding during periods of high discharge.Goudi ...

s. The Horseshoe Canyon Formation has been radiometrically dated as being between 74 and 67 million years old. It was deposited during the gradual withdrawal of the

Western Interior Seaway The Western Interior Seaway (also called the Cretaceous Seaway, the Niobraran Sea, the North American Inland Sea, and the Western Interior Sea) was a large inland sea that split the continent of North America into two landmasses. The ancient se ...

, during the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...

and

stage of the Late

period. The Horseshoe Canyon Formation is a terrestrial unit which is part of the Edmonton Group that includes the Battle Formation and the

Whitemud Member Within the earth science of geology, the Edmonton Group is a Late Cretaceous ( Campanian stage) to early Paleocene stratigraphic unit of the Western Canada Sedimentary Basin in the central Alberta plains. It was first described as the Edmonto ...

, both in

Edmonton Edmonton ( ) is the capital city of the Canadian province of Alberta. Edmonton is situated on the North Saskatchewan River and is the centre of the Edmonton Metropolitan Region, which is surrounded by Alberta's central region. The city anc ...

. The valley where dinosaurs lived included ancient meandering estuary channels, straight channels,

peat Peat (), also known as turf (), is an accumulation of partially decayed vegetation or organic matter. It is unique to natural areas called peatlands, bogs, mires, moors, or muskegs. The peatland ecosystem covers and is the most efficien ...

swamps,

river deltas A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (mo ...

s, shorelines and wetlands. Due to the changing sea levels, many different environments are represented in the Horseshoe Canyon Formation, including offshore and near-shore marine habitats and coastal habitats like

lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into '' coastal lagoons'' (or ''barrier lagoons' ...

s, and

tidal flat Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal f ...

s. The area was wet and warm with a temperate to subtropical climate. Just prior to the Campanian–Maastrichtian boundary, the mean annual temperature and precipitation in this region dropped rapidly. The dinosaurs from this formation form part of the Edmontonian land vertebrate age, and are distinct from those in the formations above and below.

Paleofauna

''Montanoceratops'' shared the paleoenvironment of the St. Mary River Formation with other dinosaurs, such as the

s ''

Anchiceratops ''Anchiceratops'' ( ) is an extinct genus of chasmosaurinae, chasmosaurine ceratopsid dinosaur that lived approximately 72 to 71 million years ago during the latter part of the Cretaceous Period (geology), Period in what is now Alberta, Canada. ' ...

'' and ''

Pachyrhinosaurus ''Pachyrhinosaurus'' (meaning in Greek "thick-nosed lizard", from ' (), thick; ' (), nose; and (), lizard) is an extinct genus of centrosaurine ceratopsid dinosaur from the Late Cretaceous period of North America. The first examples were disco ...

canadensis'', the armored

nodosaur Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, w ...

Edmontonia ''Edmontonia'' is a genus of panoplosaurin nodosaurid dinosaur from the Late Cretaceous Period. It is part of the Nodosauridae, a family within Ankylosauria. It is named after the Edmonton Formation (now the Horseshoe Canyon Formation in Canad ...

longiceps'', the duckbilled

hadrosaur Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...

Edmontosaurus ''Edmontosaurus'' ( ) (meaning "lizard from Edmonton") is a genus of hadrosaurid (duck-billed) dinosaur. It contains two known species: '' Edmontosaurus regalis'' and '' Edmontosaurus annectens''. Fossils of ''E. regalis'' have been found in ro ...

regalis'', the theropods ''

Hagryphus ''Hagryphus'' (meaning " Ha's griffin") is a monospecific genus of caenagnathid dinosaur from southern Utah that lived during the Late Cretaceous (upper Campanian stage, 75.95 Ma) in what is now the Kaiparowits Formation of the Grand Staircase� ...

'', '' Saurornitholestes'' and ''

Troodon ''Troodon'' ( ; ''Troödon'' in older sources) is a wastebasket taxon and a dubious genus of relatively small, bird-like dinosaurs known definitively from the Campanian age of the Late Cretaceous period (about 77 mya). It includes at least ...

'', and the tyrannosaurid ''

Albertosaurus ''Albertosaurus'' (; meaning "Alberta lizard") is a genus of tyrannosaurid theropod dinosaurs that lived in western North America during the Late Cretaceous Period (geology), Period, about 71 million years ago. The type species, ''A. sarc ...

'' which was likely the

apex predator An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic le ...

in its ecosystem. Vertebrates present in the St. Mary River Formation at the time of ''Montanoceratops'' included the actinopterygian fishes ''Amia fragosa'', ''

Lepisosteus ''Lepisosteus'' is a genus of gars in the family Lepisosteidae. Distribution While in the present day, ''Lepisosteus'' is only known across North America, fossil remains show it was much more widespread in the past, with specimens known from ...

'', '' Belonostomus longirostris'', '' Paralbula casei'', and ''Platacodon nanus'', the

mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Ancient Greek, Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Ma ...

'' Plioplatecarpus'', and the

diapsid Diapsids ("two arches") are a clade of sauropsids, distinguished from more primitive eureptiles by the presence of two holes, known as temporal fenestrae, in each side of their skulls. The group first appeared about three hundred million years ...

reptile ''

Champsosaurus ''Champsosaurus'' is an extinct genus of crocodile-like choristodere reptile, known from the Late Cretaceous and early Paleogene periods of North America and Europe (Campanian-Paleocene). The name ''Champsosaurus'' is thought to come from , () ...

''. A fair number of mammals lived in this region, which included ''Turgidodon russelli'', ''

Cimolestes ''Cimolestes'' (from Ancient Greek , 'chalk robber') is a genus of early eutherians with a full complement of teeth adapted for eating insects and other small animals. Paleontologists have disagreed on its relationship to other mammals, in part ...

'', '' Didelphodon'', ''

Leptalestes ''Leptalestes'' is an extinct genus of mammals in the infraclass Metatheria. It was described by B.M. Davis in 2007. A new species, ''L. toevsi'', was described from the late Cretaceous period of the United States by John P. Hunter, Ronald E. Hei ...

'', '' Cimolodon nitidus'', and '' Paracimexomys propriscus''. Non-vertebrates in this ecosystem included mollusks, the oyster ''Crassostrea wyomingensis'', the small clam ''Anomia'', and the snail ''Melania''. Plants found include the aquatic angiosperm ''Trapago angulata'', the amphibious

heterosporous Heterospory is the production of spores of two different sizes and sexes by the sporophytes of land plants. The smaller of these, the microspore, is male and the larger megaspore is female. Heterospory evolved during the Devonian period from ...

fern ''Hydropteris pinnata'', rhizomes, and taxodiaceous conifers. The paleofauna of the Horseshoe Canyon Formation featured a number of vertebrates that lived at the same time as ''Montanoceratops''. This included dinosaurs such as the

ankylosaurids Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Pal ...

'' Anodontosaurus lambei'', ''

longiceps'', ''

Euoplocephalus ''Euoplocephalus'' ( ) is a genus of very large, herbivorous ankylosaurid dinosaurs, living during the Late Cretaceous of Canada. It has only one named species, ''Euoplocephalus tutus''. The first fossil of ''Euoplocephalus'' was found in 1897 ...

tutus'', the maniraptorans '' Atrociraptor marshalli'', ''

Epichirostenotes ''Epichirostenotes'' (meaning "above '' Chirostenotes''", because it lived after the latter genus) is a genus of oviraptorosaurian dinosaur from the late Cretaceous. ''Epichirostenotes'' is known from an incomplete skeleton found in 1923 at the ...

curriei'', '' Richardoestesia gilmorei'', '' Richardoestesia isosceles'', the

troodontid Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...

'' Paronychodon lacustris'' and an unnamed species of ''

'', the

alvarezsaurid Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly ...

theropod Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...

'' Albertonykus borealis'', the

ornithomimids Ornithomimidae (meaning "bird-mimics") is a family of theropod dinosaurs which bore a superficial resemblance to modern ostriches. Ornithomimids were fast, omnivorous or herbivorous dinosaurs known mainly from the Late Cretaceous Period of Lauras ...

Dromiceiomimus ''Dromiceiomimus'' is a genus of ornithomimid theropod from the Late Cretaceous (early Maastrichtian) of Alberta, Canada. The type species, ''D. brevitertius'', is considered a synonym of '' Ornithomimus edmontonicus'' by some authors, while othe ...

brevitertius'', ''

Ornithomimus ''Ornithomimus'' (; "bird mimic") is a genus of ornithomimid dinosaurs from the Late Cretaceous Period of what is now North America. ''Ornithomimus'' was a swift bipedal theropod which fossil evidence indicates was covered in feathers, equippe ...

edmontonicus'', and an unnamed species of ''

Struthiomimus ''Struthiomimus'' (meaning "ostrich mimic", from the Greek στρούθειος/''stroutheios'' meaning "of the ostrich" and μῖμος/''mimos'' meaning "mimic" or "imitator") is a genus of ornithomimid dinosaurs from the late Cretaceous of N ...

'', the bone-head pachycephalosaurid ''

Stegoceras ''Stegoceras'' is a genus of pachycephalosaurid (dome-headed) dinosaur that lived in what is now North America during the Late Cretaceous period, about 77.5 to 74 million years ago (mya). The first specimens from Alberta, Canada, were descri ...

'', and ''

Sphaerotholus ''Sphaerotholus'' is a genus of pachycephalosaurid dinosaur from the Upper Cretaceous of the western United States and Canada. To date, three species have been described: the type species, ''S. goodwini'', from the Den-na-zin Member of the Kirtl ...

edmontonensis'', the

ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous worl ...

'' Parksosaurus'' warreni, the hadrosaurids ''

regalis'', ''

Hypacrosaurus ''Hypacrosaurus'' (meaning "near the highest lizard" reek υπο-, ''hypo-'' = less + ακρος, ''akros'', high because it was almost but not quite as large as ''Tyrannosaurus'') was a genus of hadrosaurid, duckbill dinosaur similar in app ...

altispinus'', and ''

Saurolophus ''Saurolophus'' (; meaning "lizard crest") is a genus of large hadrosaurid dinosaur from the Late Cretaceous period of Asia and North America, that lived in what is now the Horseshoe Canyon and Nemegt formations about 70 million to 68 millio ...

osborni'', the

ceratopsians Ceratopsia or Ceratopia ( or ; Ancient Greek, Greek: "horned faces") is a group of herbivore, herbivorous, beaked dinosaurs that thrived in what are now North America, Europe, and Asia, during the Cretaceous Period (geology), Period, although anc ...

ornatus'', '' Arrhinoceratops brachyops'', ''

Eotriceratops ''Eotriceratops'' (meaning "dawn three-horned face") is a genus of herbivorous ceratopsian dinosaurs which lived in the area of North America during the late Cretaceous period. The only named species is ''Eotriceratops xerinsularis''. Discovery ...

xerinsularis'', ''

canadensis'' and ''

lakustai'', the primitive

tyrannosauroid Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae as well as more basal relatives. Tyrannosauroids lived on the Laurasian supercontinent ...

'' Dryptosaurus'', and the

tyrannosaurids Tyrannosauridae (or tyrannosaurids, meaning " tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous '' Tyrannosaurus''. The exact number of ...

'' and ''

Daspletosaurus ''Daspletosaurus'' ( ; meaning "frightful lizard") is a genus of tyrannosaurid dinosaur that lived in Laramidia between about 79.5 and 74 million years ago, during the Late Cretaceous Period. The genus ''Daspletosaurus'' contains three ...

'', which were the

apex predators An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic lev ...

of this paleoenvironment. Of these, the hadrosaurs dominated in terms of sheer number and made up half of all dinosaurs who lived in this region. Vertebrates present in the Horseshoe Canyon Formation at the time of ''Montanoceratops'' included reptiles, and amphibians.

Shark Sharks are a group of elasmobranch fish characterized by a cartilaginous skeleton, five to seven gill slits on the sides of the head, and pectoral fins that are not fused to the head. Modern sharks are classified within the clade Selachi ...

s, rays, sturgeons,

bowfin The bowfin (''Amia calva'') is a bony fish, native to North America. Common names include mudfish, mud pike, dogfish, grindle, grinnel, swamp trout, and choupique. It is regarded as a relict, being the sole surviving species of the Halecomorp ...

gar Gars are members of the family Lepisosteidae, which are the only surviving members of the Ginglymodi, an ancient holosteian group of ray-finned fish, which first appeared during the Triassic, over 240 million years ago. Gars comprise seven livin ...

s and the gar-like '' Aspidorhynchus'' made up the fish fauna. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers. The saltwater plesiosaur '' Leurospondylus'' was present and freshwater environments were populated by

turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked ...

s, ''

'', and

crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest livi ...

ns like ''

Leidyosuchus ''Leidyosuchus'' (meaning " Leidy's crocodile") is an extinct genus of alligatoroid from the Late Cretaceous of Alberta. It was named in 1907 by Lawrence Lambe, and the type species is ''L. canadensis''. It is known from a number of specimens f ...

'' and ''

Stangerochampsa ''Stangerochampsa'' is an extinct genus of globidontan alligatoroid, possibly an alligatorine or a stem-caiman, from the Late Cretaceous of Alberta. It is based on RTMP.86.61.1, a skull, partial lower jaws, and partial postcranial skeleton d ...

''. Evidence has shown that

multituberculate Multituberculata (commonly known as multituberculates, named for the multiple tubercles of their teeth) is an extinct order of rodent-like mammals with a fossil record spanning over 130 million years. They first appeared in the Middle Jurassic, ...

s and the early

marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...

'' Didelphodon coyi'' were present. Vertebrate trace fossils from this region included the tracks of

theropods Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally c ...

and

ornithopods Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous worl ...

, which provide evidence that these animals were also present.Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. (2004) 861 pp. . Non-vertebrates in this ecosystem included both marine and terrestrial

invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

References

External links

{{Taxonbar, from=Q131781 Late Cretaceous dinosaurs of North America Leptoceratopsids Fossil taxa described in 1951 Paleontology in Montana Paleontology in Alberta Maastrichtian genus first appearances Maastrichtian genus extinctions Taxa named by Charles Mortram Sternberg Ornithischian genera

Description

Discovery and species

Classification

Paleobiology

Paleopathology

Paleoecology

Habitat

Paleofauna

See also

References

External links