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''Methanocaldococcus'' formerly known as ''Methanococcus'' is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
coccoid Coccoid means shaped like or resembling a coccus, that is, spherical.The noun coccoid or coccoids may refer to: * a level of organization, characterized by unicellular, non-flagellated, non-amoeboid organisms, with a definite shape, in general b ...
methanogen Methanogens are microorganisms that produce methane as a metabolic byproduct in hypoxic conditions. They are prokaryotic and belong to the domain Archaea. All known methanogens are members of the archaeal phylum Euryarchaeota. Methanogens ar ...
archaea. They are all
mesophile A mesophile is an organism that grows best in moderate temperature, neither too hot nor too cold, with an optimum growth range from . The optimum growth temperature for these organisms is 37°C. The term is mainly applied to microorganisms. Organi ...
s, except the
thermophilic A thermophile is an organism—a type of extremophile—that thrives at relatively high temperatures, between . Many thermophiles are archaea, though they can be bacteria or fungi. Thermophilic eubacteria are suggested to have been among the ear ...
''M. thermolithotrophicus'' and the
hyperthermophilic A hyperthermophile is an organism that thrives in extremely hot environments—from 60 °C (140 °F) upwards. An optimal temperature for the existence of hyperthermophiles is often above 80 °C (176 °F). Hyperthermophiles are often within the doma ...
''M. jannaschii''. The latter was discovered at the base of a “ white smoker” chimney at 21° N on the
East Pacific Rise The East Pacific Rise is a mid-ocean rise (termed an oceanic rise and not a mid-ocean ridge due to its higher rate of spreading that results in less elevation increase and more regular terrain), a divergent tectonic plate boundary located alon ...
and it was the first archaean
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding ...
to be completely sequenced, revealing many novel and
eukaryote Eukaryotes () are organisms whose cells have a nucleus. All animals, plants, fungi, and many unicellular organisms, are Eukaryotes. They belong to the group of organisms Eukaryota or Eukarya, which is one of the three domains of life. Bact ...
-like elements.


Nomenclature

The name ''Methanocaldococcus'' has Latin and Greek roots, ''methano'' for
methane Methane ( , ) is a chemical compound with the chemical formula (one carbon atom bonded to four hydrogen atoms). It is a group-14 hydride, the simplest alkane, and the main constituent of natural gas. The relative abundance of methane on Ear ...
, ''caldo'' for hot, and the Greek ''kokkos'' for the spherical shape of the cells. Overall, the name means ''spherical cell that produces methane at hot temperatures''.


Metabolism

All species in ''Methanocaldococcus'' are obligate
methanogen Methanogens are microorganisms that produce methane as a metabolic byproduct in hypoxic conditions. They are prokaryotic and belong to the domain Archaea. All known methanogens are members of the archaeal phylum Euryarchaeota. Methanogens ar ...
s. They use
hydrogen Hydrogen is the chemical element with the symbol H and atomic number 1. Hydrogen is the lightest element. At standard conditions hydrogen is a gas of diatomic molecules having the formula . It is colorless, odorless, tasteless, non-toxic ...
to reduce
carbon dioxide Carbon dioxide ( chemical formula ) is a chemical compound made up of molecules that each have one carbon atom covalently double bonded to two oxygen atoms. It is found in the gas state at room temperature. In the air, carbon dioxide is t ...
. Unlike many other species within Euryarchaeota, they cannot use
formate Formate ( IUPAC name: methanoate) is the conjugate base of formic acid. Formate is an anion () or its derivatives such as ester of formic acid. The salts and esters are generally colorless.Werner Reutemann and Heinz Kieczka "Formic Acid" in ...
,
acetate An acetate is a salt formed by the combination of acetic acid with a base (e.g. alkaline, earthy, metallic, nonmetallic or radical base). "Acetate" also describes the conjugate base or ion (specifically, the negatively charged ion called ...
, methanol or
methylamine Methylamine is an organic compound with a formula of . This colorless gas is a derivative of ammonia, but with one hydrogen atom being replaced by a methyl group. It is the simplest primary amine. Methylamine is sold as a solution in methanol, ...
s as
substrate Substrate may refer to: Physical layers *Substrate (biology), the natural environment in which an organism lives, or the surface or medium on which an organism grows or is attached ** Substrate (locomotion), the surface over which an organism lo ...
s.


Phylogeny

The currently accepted taxonomy is based on the
List of Prokaryotic names with Standing in Nomenclature List of Prokaryotic names with Standing in Nomenclature (LPSN) is an online database that maintains information on the naming and taxonomy Taxonomy is the practice and science of categorization or classification. A taxonomy (or taxonomical ...
(LPSN) and
National Center for Biotechnology Information The National Center for Biotechnology Information (NCBI) is part of the United States National Library of Medicine (NLM), a branch of the National Institutes of Health (NIH). It is approved and funded by the government of the United States. T ...
(NCBI).


See also

*
List of Archaea genera This article lists the genera of the Archaea. The currently accepted taxonomy is based on the List of Prokaryotic names with Standing in Nomenclature (LPSN) and National Center for Biotechnology Information (NCBI). Phylogeny National Center for ...
*
Methanocaldococcus sp. FS406-22 ''Methanocaldococcus'' ''sp''. FS406-22 is an archaea in the genus ''Methanocaldococcus''. It is an Anaerobic respiration, anaerobic, Piezophile, piezophilic, diazotrophic, Hyperthermophile, hyperthermophilic marine Archaea, archaeon. This strai ...


References


Further reading


Scientific journals


Scientific books

* *


Scientific databases


External links


Information on type species 'Methanocaldococcus jannaschii'''Methanocaldococcus'' at Bac''Dive'' - the Bacterial Diversity Metadatabase
Archaea genera Euryarchaeota {{Euryarchaeota-stub