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''Menefeeceratops'' (meaning "Menefee Formation horned face") is a genus of
ceratopsid Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are k ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the Menefee Formation in
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...
. It is potentially the oldest known member of the ceratopsids, as well as the
centrosaurine Centrosaurinae (from the Greek, meaning "pointed lizards") is a subfamily of ceratopsid dinosaurs, a group of large quadrupedal ornithischians. Centrosaurine fossil remains are known primarily from the northern region of Laramidia (modern day Al ...
subfamily, related to animals including '' Yehuecauhceratops'' and '' Crittendenceratops''. The
type Type may refer to: Science and technology Computing * Typing, producing text via a keyboard, typewriter, etc. * Data type, collection of values used for computations. * File type * TYPE (DOS command), a command to display contents of a file. * Ty ...
and only species is ''Menefeeceratops sealeyi,'' known from a partial, non-articulated skeleton.


Discovery and naming

The holotype, NMMNH P-25052, of ''Menefeeceratops'' was discovered by Paul Sealey in 1996 and later mentioned in a paper in 1997,. but was not given a name until it was revisited in 2021 by Sebastian Dalman, Peter Dodson, and colleagues. The holotype was collected at the NMMNH locality 3033 in
Sandoval County Sandoval County is located in the U.S. state of New Mexico. As of the 2010 United States Census, 2010 census, the population was 131,561, making it the fourth-most populous county in New Mexico. The county seat is Bernalillo, New Mexico, Bernali ...
,
New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker ...
,
USA The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country Continental United States, primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., ...
, whose rocks came from the
early Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campanian s ...
rocks of the Allison Member of the Menefee Formation. The holotype was later collected by the New Mexico Museum of Natural History with permission of the
Bureau of Land Management The Bureau of Land Management (BLM) is an agency within the United States Department of the Interior responsible for administering federal lands. Headquartered in Washington DC, and with oversight over , it governs one eighth of the country's la ...
, who owned the land the fossils were found on, and consisted of a fragmentary skull, mandible, several fragmentary vertebrae, some ribs, the left ilium, and several fragmentary limb elements. ''Menefeeceratops'' is the only Centrosaurine described from New Mexico, while many Chasmosaurines such as ''Pentaceratops'' and ''Sierraceratops'' having been described from the late Campanian and
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
. ''Menefeeceratops'' is named from the location where the fossils were found, and in honor of Paul Sealey, the research associate at the NMMNH who originally discovered them in 1996.


Description

Based on comparisons with related animals including ''
Styracosaurus ''Styracosaurus'' ( ; meaning "spiked lizard" from the Ancient Greek / "spike at the butt-end of a spear-shaft" and / "lizard") is a genus of herbivorous ceratopsian dinosaur from the Cretaceous Period (Campanian stage), about 75.5 to 74.5&nbs ...
'', ''
Vagaceratops ''Vagaceratops'' (meaning "wandering (''vagus'', Latin) horned face", in reference to its close relationship with ''Kosmoceratops'' from Utah) is a genus of herbivorous ceratopsian dinosaur. It is a chasmosaurine ceratopsian which lived during ...
'', and '' Centrosaurus'', ''Menefeeceratops'' is believed to have been approximately long in life.


Paleoenvironment

The disarticulated holotype of ''Menefeeceratops'' was collected in a greenish-gray mudstone alongside an isolated trionychid turtle costal and fragments of fossilized woods. The Menefee Formation represents a widespread alluvial floodplain and consists of
mudstone Mudstone, a type of mudrock, is a fine-grained sedimentary rock whose original constituents were clays or muds. Mudstone is distinguished from '' shale'' by its lack of fissility (parallel layering).Blatt, H., and R.J. Tracy, 1996, ''Petrology. ...
,
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility.Blatt ''et al.'' 1980, p ...
,
sandstone Sandstone is a clastic sedimentary rock composed mainly of sand-sized (0.0625 to 2 mm) silicate grains. Sandstones comprise about 20–25% of all sedimentary rocks. Most sandstone is composed of quartz or feldspar (both silicates) ...
, and
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal is formed when dea ...
. The sandstones that comprise the Menefee Formation that are fixed within carbonaceous shales of coastal swamp or
lagoon A lagoon is a shallow body of water separated from a larger body of water by a narrow landform, such as reefs, barrier islands, barrier peninsulas, or isthmuses. Lagoons are commonly divided into ''coastal lagoons'' (or ''barrier lagoons'') a ...
origin and are thought to have been created by flood tidal deltas that north and east across New Mexico and towards the retreating Western Interior Seaway. ''Menefeeceratops'' was contemporaneous with an indeterminate
ankylosaur Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. ...
, the indeterminate nodosaurid '' Invictarx'', an indeterminate
hadrosaurid Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
,Hunt, Adrian P.; Lucas, Spencer G. (1993). "Cretaceous vertebrates of New Mexico". In Lucas, S.G.; Zidek, J. (eds.). ''Dinosaurs of New Mexico''. New Mexico Museum of Natural History and Science Bulletin, 2. Albuquerque, New Mexico: New Mexico Museum of Natural History and Science. pp. 77–91. the brachylophosaurin hadrosaurid '' Ornatops'', the tyrannosaurid ''
Dynamoterror ''Dynamoterror'' (meaning "powerful terror") is a monospecific genus of tyrannosaurid dinosaur from New Mexico that lived during the Late Cretaceous (lower Campanian stage, 78 Ma) in what is now the upper Allison Member of the Menefee Formation. ...
,'' an indeterminate tyrannosaurid, and a
dromaeosaurid Dromaeosauridae () is a family of feathered theropod dinosaurs. They were generally small to medium-sized feathered carnivores that flourished in the Cretaceous Period. The name Dromaeosauridae means 'running lizards', from Greek ('), meaning ...
similar to ''
Saurornitholestes ''Saurornitholestes'' ("lizard-bird thief") is a genus of carnivorous dromaeosaurid theropod dinosaur from the late Cretaceous of Canada (Alberta) and the United States (Montana, New Mexico, Alabama, North Carolina, and South Carolina). Two spec ...
''. Non-dinosaur taxa contemporaneous with ''Menefeeceratops'' include an indeterminate crocodylian, the
alligatoroids Alligatoroidea is one of three superfamily (zoology), superfamilies of crocodylians, the other two being Crocodyloidea and Gavialoidea. Alligatoroidea evolved in the Late Cretaceous Geologic time scale#Terminology, period, and consists of the al ...
''
Brachychampsa ''Brachychampsa'' is an extinct genus of alligatoroid, possibly a basal caiman. Specimens have been reported from New Mexico, Colorado, Wyoming, Montana, North and South Dakota, New Jersey, and Saskatchewan, though only those from Montana, Utah, ...
'' and '' Deinosuchus'', an indeterminate
baenid Baenidae is an extinct family of paracryptodiran turtles known from the Early Cretaceous to Eocene of North America. While during the Early Cretaceous they are found across North America, during the Late Cretaceous they are only found in Laramidi ...
turtle Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked tu ...
, an indeterminate turtle, and an indeterminate trionychid turtle.


See also

* Timeline of ceratopsian research


References

{{Taxonbar, from=Q106797815 Centrosaurines Campanian genus first appearances Campanian genus extinctions Late Cretaceous dinosaurs of North America Fossil taxa described in 2021 Ornithischian genera