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The marbled godwit (''Limosa fedoa'') is a large migratory shorebird in the family
Scolopacidae Scolopacidae is a large family of shorebirds, or waders, which mainly includes many species known as sandpipers, but also others such as woodcocks, curlews and snipes. Most of these species eat small invertebrates picked out of the mud or soil ...
. On average, it is the largest of the four species of
godwit Godwits are a group of four large, long-billed, long-legged and strongly bird migration, migratory waders of the bird genus ''Limosa''. Their long bills allow them to probe deeply in the sand for aquatic worms and mollusca, molluscs. In their ...
.


Taxonomy

In 1750 the English naturalist George Edwards included an illustration and a description of the marbled godwit in the third volume of his ''A Natural History of Uncommon Birds''. He used the English name "The Greater American Godwit". Edwards based his hand-coloured etching on a preserved specimen that had been brought to London from the
Hudson Bay Hudson Bay, sometimes called Hudson's Bay (usually historically), is a large body of Saline water, saltwater in northeastern Canada with a surface area of . It is located north of Ontario, west of Quebec, northeast of Manitoba, and southeast o ...
area of Canada by James Isham. When in 1758 the Swedish naturalist
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
updated his ''
Systema Naturae ' (originally in Latin written ' with the Orthographic ligature, ligature æ) is one of the major works of the Sweden, Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the syste ...
'' for the tenth edition, he placed the marbled godwit with godwits and ibises in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
'' Scolopax''. Linnaeus included a brief description, coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Scolopax fedoa'' and cited Edwards' work. The marbled godwit is now placed in the genus '' Limosa'' that was introduced in 1760 by the French zoologist
Mathurin Jacques Brisson Mathurin Jacques Brisson (; 30 April 1723 – 23 June 1806) was a French zoologist and natural philosophy, natural philosopher. Brisson was born on 30 April 1723 at Fontenay-le-Comte in the Vendée department of western France. Note that page 14 ...
. The genus name ''Limosa'' is from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
and means "muddy", from ''limus'', "mud". The specific epithet ''fedoa'' may be an
Old English Old English ( or , or ), or Anglo-Saxon, is the earliest recorded form of the English language, spoken in England and southern and eastern Scotland in the Early Middle Ages. It developed from the languages brought to Great Britain by Anglo-S ...
name for a godwit. The word was mentioned by the English naturalist William Turner in 1544. Two
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised: *''L. f. beringiae'' Gibson & Kessel, 1989 – breeds in Alaska and winters in the west United States. Averages shorter legs and bill than nominate subspecies. *''L. f. fedoa'' (
Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
, 1758) – breeds in central, south-central Canada, and the north-central United States, winters in southern United States to northwest South America. A disjunct breeding population breeds in
James Bay James Bay (, ; ) is a large body of water located on the southern end of Hudson Bay in Canada. It borders the provinces of Quebec and Ontario, and is politically part of Nunavut. Its largest island is Akimiski Island. Numerous waterways of the ...
.


Description

The total length is , including a large bill of , and wingspan is . Body mass can vary from . The average weight of 40 males was and that of 45 females was . Bill length is from . Among all the members of the sandpiper family, only the
curlew The curlews () are a group of nine species of birds in the genus ''Numenius'', characterised by their long, slender, downcurved bills and mottled brown plumage. The English name is imitative of the Eurasian curlew's call, but may have been infl ...
s attain sizes that significantly exceed this species.''CRC Handbook of Avian Body Masses'' by John B. Dunning Jr. (Editor). CRC Press (1992), . Adults have long blue-grey legs and a very long pink bill with a slight upward curve and dark at the tip. The long neck, breast and belly are pale brown with dark bars on the breast and flanks. The back is mottled and dark. They show cinnamon wing linings in flight.


Distribution and habitat

Marbled godwits breed in three distinct areas with their own unique route. The vast majority occur in mid-continental North America, followed by eastern Canada and the Alaska Peninsula, USA. In addition, the largest winter ranges are the Atlantic, Pacific and Gulf coasts of the US and Mexico. Godwits breeding in the western USA and Canada follow a route through the Utah stopover site, with a final arrival in the winter sites of Mexico and the Caribbean. Species breeding in eastern Canada migrate across the US, and stopover at sites along the Gulf of California and Mexico. Furthermore, those breeding in North and South Dakota winter in coastal Georgia. The Bear River Migratory Bird Refuge located at Great Salt Lake in Utah (USA), is one of the most popular stopover sites for godwits in the spring and fall.


Behavior and ecology


Breeding

They nest on the ground, usually in short grass.


Food and feeding

These birds forage by probing on mudflats, in
marsh In ecology, a marsh is a wetland that is dominated by herbaceous plants rather than by woody plants.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p More in genera ...
es, or at the beach (see picture below). When the tide is out, they eat. In short grass, they may pick up insects by sight. They mainly eat insects and
crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...
s, but also eat parts of aquatic plants. When the tide is in, they roost. They often sleep by standing on one leg and tucking their bill into their body (see picture below).


Conservation

Their numbers were reduced by hunting at the end of the 19th century. Although they had recovered somewhat since that time, their population has declined in recent times as suitable habitat is used for farming.


Gallery

File:Marbled Godwit Northern California.jpg, Feeding on mudflat at low tide, Elkhorn Slough File:MarbledGodwitsFeeding.jpg, Marbled godwits feeding,
Point Reyes National Seashore Point Reyes National Seashore is a park preserve located on the Point Reyes, Point Reyes Peninsula in Marin County, California. As a national seashore, it is maintained by the US United States National Park Service, National Park Service as an ...
, California File:Marbled godwit Sandiego.jpg, Marbled godwit landing
San Diego San Diego ( , ) is a city on the Pacific coast of Southern California, adjacent to the Mexico–United States border. With a population of over 1.4 million, it is the List of United States cities by population, eighth-most populous city in t ...
CA File:Sea foam at Ocean Beach in San Francisco -1 on 3-25-11.jpg, Marbled godwit flying near sea foam at Ocean Beach,
San Francisco San Francisco, officially the City and County of San Francisco, is a commercial, Financial District, San Francisco, financial, and Culture of San Francisco, cultural center of Northern California. With a population of 827,526 residents as of ...
File:Limosa fedoa flight.jpg, In flight File:7Z1E8737.jpg, Flock File:Marbled godwits napping on the beach.jpg, Sleeping at Balboa Peninsula, Newport Beach, California. File:Limosa fedoa (Marbled Godwit) 1APR2017.jpg, Bodega Bay, California


References


External links


Marbled Godwit Species Accounty
- Cornell Lab of Ornithology

- USGS Patuxent Bird Identification InfoCenter * * {{Taxonbar, from=Q844714 Birds described in 1758 Birds of the Caribbean Birds of the Dominican Republic Limosa Native birds of the Canadian Prairies Native birds of the Plains-Midwest (United States) Shorebirds Taxa named by Carl Linnaeus