''Mansourasaurus'' ("
Mansoura lizard
Lizards are a widespread group of squamate reptiles, with over 7,000 species, ranging across all continents except Antarctica, as well as most oceanic island chains. The group is paraphyletic since it excludes the snakes and Amphisbaenia alt ...
") is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of
herbivorous lithostrotian
sauropod
Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their bo ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
from the
Quseir Formation
The Quseir Formation is a geological formation in the vicinity of the Kharga Oasis in Egypt. of
Egypt
Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a transcontinental country spanning the northeast corner of Africa and southwest corner of Asia via a land bridge formed by the Sinai Peninsula. It is bordered by the Mediter ...
. The
type and only species is ''Mansourasaurus shahinae''.
The discovery of ''Mansourasaurus'' was considered quite significant by
paleontologist
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
s, because very few
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
sauropod remains had been found in Africa where the rocky
strata
In geology and related fields, a stratum ( : strata) is a layer of rock or sediment characterized by certain lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by visible surfaces known as ei ...
that preserve remains elsewhere and produce rich fossil beds were typically not found exposed at or near ground level.
Discovery and naming
Hesham Sallam
Hesham Sallam ( ar, هشام سلام; born 1975) is an Egyptian paleontologist and the founder of the Mansoura University Vertebrate Paleontology Center (MUVP-C), the first vertebrate paleontology program in the Middle East. He works as an assoc ...
, a paleontologist at
Mansoura University, together with a team of students discovered a sauropod skeleton in the
Dakhla Oasis in Egypt's Western Desert.
In 2016, it was reported that over thirty dinosaur specimens had been excavated, among them titanosaurian sauropods.
[Iman El-Dawoudi, Patrick M O'Connor, Mahmoud Kora and Joseph J W Sertich, 2016, "NEW DINOSAUR REMAINS FROM THE CAMPANIAN QUSEIR FORMATION, WESTERN DESERT, EGYPT", ''SVP October 2016, Program and Abstracts'', p 129]
Based on this skeleton, the type species ''Mansourasaurus shahinae'' was named and described in January 2018, by Hesham M. Sallam, Eric Gorscak, Patrick M. O'Connor, Iman A. El-Dawoudi, Sanaa El-Sayed, Sara Saber, Mahmoud A. Kora, Joseph J. W. Sertich, Erik R. Seiffert and Matthew C. Lamanna. The generic name refers to the Mansoura University. The
specific name Specific name may refer to:
* in Database management systems, a system-assigned name that is unique within a particular database
In taxonomy, either of these two meanings, each with its own set of rules:
* Specific name (botany), the two-part (bino ...
honours Mona Shahin, one of the founders of the Mansoura University Vertebrate Paleontology Center.
The ''Mansourasaurus''
specimen
Specimen may refer to:
Science and technology
* Sample (material), a limited quantity of something which is intended to be similar to and represent a larger amount
* Biological specimen or biospecimen, an organic specimen held by a biorepository ...
described in 2018 is its
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
, MUVP 200, discovered in a layer of the Quseir Formation dating from the late
Campanian, about seventy-three million years old. It consists of a partial skeleton with skull and lower jaws.
It contains a fragment of the skull roof, a part of the lower
braincase
In human anatomy, the neurocranium, also known as the braincase, brainpan, or brain-pan is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calvaria or skul ...
, the dentaries of the lower jaws, three neck vertebrae, two back vertebrae, eight ribs, the right scapula, the right
coracoid, both humeri, a radius, a third
metacarpal, three
metatarsals, and parts of
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinct amp ...
s. The skeleton was found on a surface of four by three metres. It was not articulated. The authors concluded that the holotype is a juvenile specimen, because the bones of its shoulder girdle had not yet fused. An
ulna
The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...
, specimen MUVP 201, found at twenty metres distance from the skeleton, was not referred to the species as it seemed somewhat too large for the holotype individual and a general connection to the species could not be proven.
Description
The—not fully-grown—holotype individual was about long.
It probably weighed about 5,000 kg, approximately the same as a
bull African elephant.
The describing authors indicated a number of distinguishing traits. These are
autapomorphies, unique derived characters. Each lower jaw dentary bears ten teeth. Where the dentaries touch each other, at the front of the lower jaws, they possess a common "chin", equalling a third of the front height. The horizontal groove in the inner side of the dentary, the ''fossa Meckeliana'', largely opens to below. The anterior middle neck vertebrae are pierced by a
in the rear side. In at least one anterior middle neck vertebra the parapophysis, the process which bears the facet for the lower rib head, has a horizontal length equal to the vertebral centrum as a whole. With some anterior neck vertebrae, the bone web between the heads of the neck rib is pierced by a foramen. The lower end of the radius has a transverse width four times larger than the width measured from the front to the rear.
Phylogeny
''Mansourasaurus'' was placed in the
Titanosauria in a derived position as a
sister species of ''
Lohuecotitan
''Lohuecotitan'' is an extinct genus of titanosaurian sauropod dinosaur which lived during the Late Cretaceous in Spain. The only species known in the genus is ''Lohuecotitan pandafilandi'', described and named in 2016..
Discovery and naming
T ...
''. A
cladistic analysis showed it to belong to a
clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, ...
of otherwise largely Eurasian sauropods, also including ''
Ampelosaurus'', ''
Lirainosaurus'', ''
Nemegtosaurus'', ''
Opisthocoelicaudia'' and ''
Paludititan
''Paludititan'' is a genus of titanosaurian sauropod dinosaur which lived in the area of present Romania during the Late Cretaceous. It existed in the island ecosystem known as Hațeg Island.
Discovery and naming
In 2002, a Belgian-Romanian exp ...
'', more or less contemporaneous forms. Hypotheses about relationships between
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
African and Eurasian sauropods had been hard to test, because very few of their remains had been found in Africa. ''Mansourasaurus'' represents the best-known continental African (i.e. excluding
Madagascar
Madagascar (; mg, Madagasikara, ), officially the Republic of Madagascar ( mg, Repoblikan'i Madagasikara, links=no, ; french: République de Madagascar), is an island country in the Indian Ocean, approximately off the coast of East Africa ...
) titanosaur of the
Upper Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
from the time period after the
Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the s ...
. Its existence would show that the continent was far less isolated from the various Eurasian landmasses than had been assumed. The ancestors of ''Mansourasaurus'' would have reached Africa from Europe.
See also
*
2018 in paleontology
Flora Plants
Fungi
Cnidarians
Research
* New three dimensionally phosphatized microfossils of coronate scyphozoan '' Qinscyphus necopinus'', including a new type of fossil embryo, are described from the Cambrian (Fortunian) Kuanchuanpu F ...
References
{{Taxonbar, from=Q47518933
Lithostrotians
Late Cretaceous dinosaurs of Africa
Campanian life
Fossils of Egypt
Fossil taxa described in 2018