Malesia is a
biogeographical
Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, i ...
region straddling the
Equator
The equator is a circle of latitude, about in circumference, that divides Earth into the Northern and Southern hemispheres. It is an imaginary line located at 0 degrees latitude, halfway between the North and South poles. The term can als ...
and the boundaries of the
Indomalayan and
Australasian realms, and also a
phytogeographical floristic region in the
Paleotropical Kingdom. It has been given different definitions. The
World Geographical Scheme for Recording Plant Distributions split off
Papuasia
Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia.
It comprises the ...
in its 2001 version.
Floristic province
Malesia was first identified as a floristic region that included the
Malay Peninsula
The Malay Peninsula ( Malay: ''Semenanjung Tanah Melayu'') is a peninsula in Mainland Southeast Asia. The landmass runs approximately north–south, and at its terminus, it is the southernmost point of the Asian continental mainland. The are ...
, the
Malay Archipelago,
New Guinea
New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torr ...
, and the
Bismarck Archipelago
The Bismarck Archipelago (, ) is a group of islands off the northeastern coast of New Guinea in the western Pacific Ocean and is part of the Islands Region of Papua New Guinea. Its area is about 50,000 square km.
History
The first inhabitants o ...
, based on a shared tropical flora derived mostly from
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
but also with numerous elements of the
Antarctic flora, including many species in the southern
conifer
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...
families
Podocarpaceae and
Araucariaceae
Araucariaceae – also known as araucarians – is an extremely ancient family of coniferous trees. The family achieved its maximum diversity during the Jurassic and Cretaceous periods and the early Cenozoic, when it was distributed almost world ...
. The floristic region overlaps four distinct
mammalian
faunal regions
Zoogeography is the branch of the science of biogeography that is concerned with geographic distribution (present and past) of animal species.
As a multifaceted field of study, zoogeography incorporates methods of molecular biology, genetics, mor ...
.
The first edition of the
World Geographical Scheme for Recording Plant Distributions (WGSRPD) used this definition, but in the second edition of 2001, New Guinea and the Bismarck Archipelago were removed from Malesia and were united with the
Solomon Islands
Solomon Islands is an island country consisting of six major islands and over 900 smaller islands in Oceania, to the east of Papua New Guinea and north-west of Vanuatu. It has a land area of , and a population of approx. 700,000. Its ca ...
, previously placed in the WGSRPD's Southwestern Pacific region, being placed into a new region,
Papuasia
Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia.
It comprises the ...
.
Sundaland
The western part of Malesia, which includes the Malay Peninsula and the islands of
Sumatra,
Java
Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
,
Bali
Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and Nu ...
, and
Borneo
Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
, shares the large mammal fauna of Asia and is known as
Sundaland
Sundaland (also called Sundaica or the Sundaic region) is a biogeographical region of South-eastern Asia corresponding to a larger landmass that was exposed throughout the last 2.6 million years during periods when sea levels were lower. It ...
. These islands are on Asia's relatively shallow
continental shelf, and were linked to Asia during the
ice ages, when sea levels were lower. The eastern edge of Sundaland is the
Wallace line, named after
Alfred Russel Wallace, the nineteenth-century British naturalist who noted the difference in fauna between islands on either side of the line.
Dipterocarps are predominant trees in the lowland forests of Sundaland. Sundaland has the greatest diversity of Dipterocarp species, with 10 to 14 native genera and approximately 450 native species, including approximately 267 species on Borneo, 155 on the Malay Peninsula, and 106 on Sumatra.
[G. Maury-Lechon and L. Curtet (1998). "Biogeography and Evolutionary Systematics of Dipterocarpaceae". in ''A Review of Dipterocarps: Taxonomy, ecology and silviculture''. Simmathiri Appanah & Jennifer M. Turnbull, eds. Center for International Forestry Research, 1998, Bogor Indonesia.]
Philippines
Most of the
Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
were never connected to the Asian mainland, and have a largely Asian-derived flora, and a distinct mammalian fauna.
The Philippines have approximately 50 species of Dipterocarps in 11 genera.
[
]
Wallacea
The islands between Sundaland and New Guinea, called Wallacea, were never linked to the neighboring continents, and have a flora and fauna that include Indomalayan and Australasian elements.
Dipterocarps, which are dominant in Sundaland, are less common in Wallacea, with only 13 species in 4 genera.[
]
New Guinea and the Bismarck Archipelago (Sahulland)
Together with the Solomon Islands, New Guinea and the Bismarck Archipelago are placed in Papuasia
Papuasia is a Level 2 botanical region defined in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It lies in the southwestern Pacific Ocean, in the Melanesia ecoregion of Oceania and Tropical Asia.
It comprises the ...
rather than Malesia in the second version of the WGSRPD. The eastern end of the earlier definition of Malesia, which includes New Guinea and the Aru Islands of eastern Indonesia, is linked to Australia by a shallow continental shelf, and shares many marsupial
Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
mammal and bird taxa
In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
with Australia. New Guinea also has many additional elements of the Antarctic flora, including southern beech (''Nothofagus
''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere in southern South America (Chile, Argentina) and Australasia (east and southeast Australia, New Zealand, New Gu ...
'') and eucalypt
Eucalypt is a descriptive name for woody plants with capsule fruiting bodies belonging to seven closely related genera (of the tribe Eucalypteae) found across Australasia:
''Eucalyptus'', '' Corymbia'', '' Angophora'', '' Stockwellia'', '' Allo ...
s. New Guinea has the highest mountains in Malesia and Papuasia, and vegetation ranges from tropical lowland forest to tundra.
Assembly and origins of the Malesian flora
Major contributions to rainforest
Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainfo ...
assembly have come from floristic elements which were carried on the Indian Plate
The Indian Plate (or India Plate) is a minor tectonic plate straddling the equator in the Eastern Hemisphere. Originally a part of the ancient continent of Gondwana, the Indian Plate broke away from the other fragments of Gondwana , began m ...
and montane elements which have come from the Australian Plate ( Sahul). The Sahul component is now understood to include substantial two-way exchanges with Sunda inclusive of lowland taxa. Evidence for the relative contributions of the great Asiatic floristic interchanges (GAFIs) with India and Sahul, respectively, to the flora of Malesia comes from contemporary lineage distributions, the fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
record, time-calibrated phylogenies, functional traits, and the spatial structure of genetic diversity. Functional trait and biome conservatism are noted features of montane austral lineages from Sahul (e.g., diverse Podocarpaceae), whereas the abundance and diversity of lowland lineages, including groups such as Syzygium (Myrtaceae
Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All spe ...
) and the Asian dipterocarps (Dipterocarpoideae), reflect a less well understood combination of dispersal, ecology, and adaptive radiations. Thus, Malesian rainforest assembly has been shaped by sharply contrasting evolutionary origins and biogeographic histories.
See also
* Nusantara
* Phytochorion - floristic regions and provinces
* Coral Triangle
* Wallacea
References
External links
*
*
{{floristic regions
Australasian realm
Indomalayan realm
Paleotropical flora
Floristic regions