Macrofossils, also known as megafossils, are the preserved remnants of
organic beings and their activities that are large enough to be visible without a
microscope
A microscope () is a laboratory equipment, laboratory instrument used to examine objects that are too small to be seen by the naked eye. Microscopy is the science of investigating small objects and structures using a microscope. Microscopic ...
. The term ''macrofossil'' stands in opposition to the term
microfossil. Microfossils, by contrast, require substantial magnification for evaluation by
fossil
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
-hunters or professional
paleontologist
Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
s. As a result, most fossils observed in the
field and most specimens are macrofossils. Macrofossils come in many varieties and form in various ways depending on their environment and what is being fossilized including plant, fungi and animal remnants.
Historical background
Etymology
The origins of the term macrofossil dates back to 1937 and was first used by G. D. Hanna, a US paleontologist and
malacologist
Malacology, from Ancient Greek μαλακός (''malakós''), meaning "soft", and λόγος (''lógos''), meaning "study", is the branch of invertebrate zoology that deals with the study of the Mollusca (molluscs or mollusks), the second-largest ...
.
Scientific thought
Macrofossils have been subject to analysis since ancient times. Several Greek philosophers used fossilized shellfish to theorize the land was once covered by oceans. Macrofossils of body parts have been used to reconstruct extinct animals and provide basis for
evolutionary lineages.
Georges Cuvier
Jean Léopold Nicolas Frédéric, baron Cuvier (23 August 1769 – 13 May 1832), known as Georges Cuvier (; ), was a French natural history, naturalist and zoology, zoologist, sometimes referred to as the "founding father of paleontology". Cuv ...
, a
naturalist
Natural history is a domain of inquiry involving organisms, including animals, fungi, and plants, in their natural environment, leaning more towards observational than experimental methods of study. A person who studies natural history is cal ...
,
zoologist
Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...
and the founding father of paleontology, along with other natural thinkers during the
Age of Reason
The Age of Enlightenment (also the Age of Reason and the Enlightenment) was a European intellectual and philosophical movement active from the late 17th to early 19th century. Chiefly valuing knowledge gained through rationalism and empiric ...
presented fossils as integral evidence in the changing views of
natural philosophy
Natural philosophy or philosophy of nature (from Latin ''philosophia naturalis'') is the philosophical study of physics, that is, nature and the physical universe, while ignoring any supernatural influence. It was dominant before the develop ...
at the time from the application of new mathematics and other emerging branches of science. This brought about the acceptance of paleontology as a true scientific discipline. This also greatly affected the development of
geology
Geology (). is a branch of natural science concerned with the Earth and other astronomical objects, the rocks of which they are composed, and the processes by which they change over time. Modern geology significantly overlaps all other Earth ...
and
stratigraphy
Stratigraphy is a branch of geology concerned with the study of rock layers (strata) and layering (stratification). It is primarily used in the study of sedimentary and layered volcanic rocks.
Stratigraphy has three related subfields: lithost ...
as new schools of science. This led to other scientific thinkers like
Charles Darwin
Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English Natural history#Before 1900, naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all speci ...
, a naturalist,
biologist
A biologist is a scientist who conducts research in biology. Biologists are interested in studying life on Earth, whether it is an individual Cell (biology), cell, a multicellular organism, or a Community (ecology), community of Biological inter ...
and
geologist
A geologist is a scientist who studies the structure, composition, and History of Earth, history of Earth. Geologists incorporate techniques from physics, chemistry, biology, mathematics, and geography to perform research in the Field research, ...
, to begin using macrofossils of evidence in a new scientific discipline called
evolutionary biology
Evolutionary biology is the subfield of biology that studies the evolutionary processes such as natural selection, common descent, and speciation that produced the diversity of life on Earth. In the 1930s, the discipline of evolutionary biolo ...
.
Varieties
Plant macrofossils
Plant
Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they are predominantly Photosynthesis, photosynthetic. This means that they obtain their energy from sunlight, using chloroplasts derived from endosymbiosis with c ...
macrofossils include
leaf
A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leav ...
,
needle,
cone
In geometry, a cone is a three-dimensional figure that tapers smoothly from a flat base (typically a circle) to a point not contained in the base, called the '' apex'' or '' vertex''.
A cone is formed by a set of line segments, half-lines ...
, and
stem
Stem or STEM most commonly refers to:
* Plant stem, a structural axis of a vascular plant
* Stem group
* Science, technology, engineering, and mathematics
Stem or STEM can also refer to:
Language and writing
* Word stem, part of a word respon ...
debris; and can be used to identify types of plants formerly growing in the area. Such
botanical
Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
macrofossil data provide a valuable complement to
pollen
Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
and
fauna
Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...
l data that can be used to reconstruct the
prehistoric
Prehistory, also called pre-literary history, is the period of human history between the first known use of stone tools by hominins million years ago and the beginning of recorded history with the invention of writing systems. The use o ...
terrestrial environment.
Algal
Algae ( , ; : alga ) is an informal term for any organisms of a large and diverse group of photosynthetic organisms that are not plants, and includes species from multiple distinct clades. Such organisms range from unicellular microalgae, s ...
macrofossils (for instance, brown
kelp
Kelps are large brown algae or seaweeds that make up the order (biology), order Laminariales. There are about 30 different genus, genera. Despite its appearance and use of photosynthesis in chloroplasts, kelp is technically not a plant but a str ...
,
sea lettuce
The sea lettuces comprise the genus ''Ulva'', a group of edible green algae that is widely distributed along the coasts of the world's oceans. The type species within the genus ''Ulva'' is ''Ulva lactuca'', wikt:lactuca, ''lactuca'' being Latin ...
and large
stromatolite
Stromatolites ( ) or stromatoliths () are layered Sedimentary rock, sedimentary formation of rocks, formations (microbialite) that are created mainly by Photosynthesis, photosynthetic microorganisms such as cyanobacteria, sulfate-reducing micr ...
s) are increasingly used to analyze prehistoric
marine and
aquatic ecosystem
An aquatic ecosystem is an ecosystem found in and around a body of water, in contrast to land-based terrestrial ecosystems. Aquatic ecosystems contain communities of organisms—aquatic life—that are dependent on each other and on their environ ...
s. The study of these fossils is called
paleobotany
Paleobotany or palaeobotany, also known as paleophytology, is the branch of botany dealing with the recovery and identification of plant fossils from geological contexts, and their use for the biological reconstruction of past environments ( pal ...
. Plant macrofossils are increasingly being used along with pollen microfossils to reconstruct past climates.
Vertebrate macrofossils
Vertebrate
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain.
The vertebrates make up the subphylum Vertebra ...
macrofossils include the
teeth
A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tear ...
,
skull
The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate.
In the human, the skull comprises two prominent ...
s, and
bone
A bone is a rigid organ that constitutes part of the skeleton in most vertebrate animals. Bones protect the various other organs of the body, produce red and white blood cells, store minerals, provide structure and support for the body, ...
s. Vertebrate macrofossils are used to reconstruct extinct animals, determine their behaviors and how they evolved. Well preserved body fossils are of particular use in helping to reassemble extinct animal body structures.
Invertebrate macrofossils
Invertebrate
Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
macrofossils include remains such as
shells,
tests
Test(s), testing, or TEST may refer to:
* Test (assessment), an educational assessment intended to measure the respondents' knowledge or other abilities
Arts and entertainment
* ''Test'' (2013 film), an American film
* ''Test'' (2014 film) ...
,
faunal armor, and
exoskeleton
An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the internal organs, in contrast to an internal endoskeleton (e.g. human skeleton, that ...
s. The bodies of small and soft bodied invertebrates rarely fossilize. Meanwhile, large and hard bodied invertebrates more commonly fossilize. The study of these fossils is a subdivision of paleontology called
invertebrate paleontology
Invertebrate paleontology (also spelled invertebrate palaeontology) is sometimes described as invertebrate paleozoology or invertebrate paleobiology.
Whether it is considered to be a subfield of paleontology, paleozoology, or paleobiology, this d ...
.
Fungi macrofossils
Fungi
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one ...
macrofossils include
fungal bodies and
filaments. Most fungi fossils are found in
amber
Amber is fossilized tree resin. Examples of it have been appreciated for its color and natural beauty since the Neolithic times, and worked as a gemstone since antiquity."Amber" (2004). In Maxine N. Lurie and Marc Mappen (eds.) ''Encyclopedia ...
. Fungi fossils are under-researched compared to animal and plant fossils. Macrofossils of fungi are rarer and are more commonly found as microfossils. The study of these fossils is called
paleomycology.
Trace macrofossils
Trace macrofossils are fossilized evidence of animal
behavior
Behavior (American English) or behaviour (British English) is the range of actions of Individual, individuals, organisms, systems or Artificial intelligence, artificial entities in some environment. These systems can include other systems or or ...
. These may include
tracks,
burrow
file:Chipmunk-burrow (exits).jpg, An eastern chipmunk at the entrance of its burrow
A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of Animal lo ...
s and dung (
coprolite
A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
s). Trace fossils are the primary type of fossils used by paleontologist to reconstruct extinct animal behaviors. The study of these fossils is called
Ichnology.
Chemo macrofossils
Chemo-fossils are the fossilized chemical remains of an organism. For example
coal
Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen.
Coal i ...
is the fossilized chemical remains of extinct plant matter.
Image gallery
Petrified Araucaria cone from patagonia-Edit3.jpg, Jurassic
The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
Petrified cone of ''Araucaria'' sp. from Patagonia
Patagonia () is a geographical region that includes parts of Argentina and Chile at the southern end of South America. The region includes the southern section of the Andes mountain chain with lakes, fjords, temperate rainforests, and glaciers ...
, Argentina
Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
.
Cockerellites liops 18" layer img2.jpg, Eocene fossil fish ''Priscacara liops'' from the Green River Formation
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River (Colorado River), Green River in Colorado, Wyoming, and Utah. The sedimen ...
of Utah.
RudistCretaceousUAE.jpg, Rudist bivalves from the Cretaceous of the Omani Mountains, United Arab Emirates. Scale bar is 10 mm.
Palaeopascichnus_CU21.png, The enigmatic Ediacaran
The Ediacaran ( ) is a geological period of the Neoproterozoic geologic era, Era that spans 96 million years from the end of the Cryogenian Period at 635 Million years ago, Mya to the beginning of the Cambrian Period at 538.8 Mya. It is the last ...
lifeform known as Palaeopascichnus is known as a ''Macrofossil''.
References
{{Reflist
Fossils