Lycaena Virgaureae
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The scarce copper (''Lycaena virgaureae'') is a
butterfly Butterflies are insects in the macrolepidopteran clade Rhopalocera from the Order (biology), order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The ...
of the family
Lycaenidae Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...
(copper or gossamer-winged butterflies).


Appearance

The lower surfaces of the back wings are yellowish and have only a few black dots; there are characteristic white marks in the immediate vicinity of these. The species exhibits one kind of
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
: male butterflies are colored bright gold-red on the upper side of wing, while the females have broader orange wings with a dark design.


Life cycle

A generation appears from mid-July to mid-September. Eggs are laid on dried-out plant parts, for example on dry
sorrel Sorrel (''Rumex acetosa''), also called common sorrel or garden sorrel, is a perennial herbaceous plant in the family Polygonaceae. Other names for sorrel include spinach dock and narrow-leaved dock ('dock' being a common name for the genus '' ...
stems. The eggs are white in color and somewhat larger than those of other ''Lycaena'' species. The caterpillars are green and nocturnal and eat sorrel. ''Lycaena virgaureae'' is the only species of this genus whose eggs last over the winter. The butterflies feed on blossoms of such plants as the
ground-elder ''Aegopodium podagraria'', commonly called ground elder, is a species of flowering plant in the carrot family Apiaceae that grows in shady places. The name "ground elder" comes from the superficial similarity of its leaves and flowers to those o ...
, ''
Eupatorium ''Eupatorium'' is a genus of flowering plants in the family Asteraceae, containing from 36 to 60 species depending on the classification system. Most are herbaceous perennials growing to tall. A few are shrubs. The genus is native to temperate ...
'', ''
Valeriana ''Valeriana'' is a genus of flowering plants in the family Caprifoliaceae, members of which may by commonly known as valerians. It contains many species, including the garden valerian, ''Valeriana officinalis''. Species are native to all continen ...
'', and
burnet saxifrage ''Pimpinella saxifraga'', known as burnet-saxifrage, solidstem burnet saxifrage, lesser burnet is a plant species in the family Apiaceae, a native of the British Isles and temperate Europe and Western Asia. It is neither a burnet, which its l ...
.


Habitat

''Lycaena virgaureae'' lives preferably on flower-rich, dry and damp meadows in
Central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the area' ...
. In
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = ''Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , i ...
it is found in the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to C ...
and in the
Cantabrian Mountains , etymology=Named after the Cantabri , photo=Cordillera Cantábrica vista desde el Castro Valnera.jpg , photo_caption=Cantabrian Mountains parallel to the Cantabrian Sea seen from Castro Valnera in an east-west direction. In the background, th ...
. In
southern France Southern France, also known as the South of France or colloquially in French language, French as , is a defined geographical area consisting of the regions of France that border the Atlantic Ocean south of the Marais Poitevin,Louis Papy, ''Le midi ...
it is found in the
Massif Central The (; oc, Massís Central, ; literally ''"Central Massif"'') is a highland region in south-central France, consisting of mountains and plateaus. It covers about 15% of mainland France. Subject to volcanism that has subsided in the last 10,00 ...
. Its area of distribution reaches the
Arctic Circle The Arctic Circle is one of the two polar circles, and the most northerly of the five major circles of latitude as shown on maps of Earth. Its southern equivalent is the Antarctic Circle. The Arctic Circle marks the southernmost latitude at w ...
in
Fennoscandia __NOTOC__ Fennoscandia (Finnish language, Finnish, Swedish language, Swedish and no, Fennoskandia, nocat=1; russian: Фенноскандия, Fennoskandiya) or the Fennoscandian Peninsula is the geographical peninsula in Europe, which includes ...
in the north; in the south it reaches north and northwest
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
. In
Belgium Belgium, ; french: Belgique ; german: Belgien officially the Kingdom of Belgium, is a country in Northwestern Europe. The country is bordered by the Netherlands to the north, Germany to the east, Luxembourg to the southeast, France to th ...
there are a few threatened populations in the south of the country. It is not found, however, on the
British Isles The British Isles are a group of islands in the North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner and Outer Hebrides, the Northern Isles, ...
or the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
. It can be found as far east as
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million, ...
. The scarce copper is found at elevations of 1,000 to 2,000 meters, although in Armenia it inhabits meadows up to 2,700 meters above sea level. There have been several scientific debates about the Scarce Copper presence in Britain during the 19th century.


Subspecies

* ''Lycaena virgaureae montanus'' Meyer-Dür, 1851 Found in the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
in
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
, and
Austria Austria, , bar, Östareich officially the Republic of Austria, is a country in the southern part of Central Europe, lying in the Eastern Alps. It is a federation of nine states, one of which is the capital, Vienna, the most populous ...
at elevations of . * ''Lycaena virgaureae miegii'' Vogel, 1857 North and central Spain at elevations of . * ''Lycaena virgaureae armeniaca'' (A. Bang-Haas, 1906) Endemic of Armenia, occupies elevation range .


Description from Seitz

Male above red-golden, with a narrow black margin, at the proximal side of which there are dark dots on the hindwing: female cinnabar-red, spotted with black, the hindwing partly dusted with black. Underside leather-yellow, more sparsely spotted with black, before the outer third of the hind- wing pale dots, which are occasionally united in a white chain. The anal area dusted with red. The species occurs from the Atlantic coasts throughout Europe to East- Siberia and from the coast of the North Sea to the Mediterranean, but is absent from Great Britain and Japan. — In Lapland flies a small form, ''oranula'' Frr. (76 b), which is otherwise very similar to the name-typical form. — ''estonica'' Huene (76 b), from the Baltic provinces and eastern Russia, exactly resembles the preceding in size and shape, but has a broader black margin. —'' virgaureola'' Stgr. (76 b) is in size like ''virgaureae'', but the upperside of the male is as in ''estonica''; beneath the more reddish disc of the forewing contrasts with the more yellow hindwing and the white discal stripe of the latter is absent or reduced; from northern Central Asia, Dauria, Mongolia, and as similar aberration in the Swiss Alps. — males from the Apennines, in which the upperside is deeper red-golden and the base of the hindwing more densely dusted with dark, are ''apennina'' Calb. The females the contrary have the ground-colour paler and are less dusted with dark than many Central European specimens, the underside being hghter. According to Fallou the males from the Pyrenees are also deeper red. - On the other hand, the specimens from Mersina and the neighbouring Taurus Mts. have a bright light golden-red upperside in both sexes, the black margin being narrowed in the male. This is aureomicans Heyne. — Specimens with the underside of the hindwing strongly dusted with grey, the upperside of the female more-over having a brown-grey tint on account of the dark dusting on the golden ground, occur in many alpine districts with the ordinary form, being especially plentiful and well marked in the Alps of Valais; all such individuals are united as ab. ''zermattensis'' Fall. (76 a). It is hardly possible to decide from Fallou's very long description of this form which are the real characteristics of his aberration, so that strictly speaking not all the specimens standing in collections under this name belong here, particularly not all the specimens from the Valais; in fact only those individuals are true ab. zermattensis which, like ''phlaeas caeruleopunctata'', have a chain of white dots before the outer third of the hindwing, as shown in the figure given in ''Ann. Soc. Ent. Fr.'' vol. V. pl. 2. Besides the modification of the colour of the upperside into sepia-brown, both sexes differ on the underside in the forewing bearing a broad dark margin and the hindwing being washed with fuscous, ab. ''seriata'' Fruhst. are ''zermattensis'' females which have a discal row of white dots on the hindwing proximally to the row of black spots, ab. ''fredegunda'' Fruhst. bears, besides those dots, a row of submarginal spots on the forewing beneath. ore vars describedhe egg is semiglobular, grey-green, with a network of polygons, laid singly or several together on the stalk of the foot-plant. Larva green with a yellow stripe on back and at the sides, brownish head and brownish legs; at first glassy, transparent, later yellow on the back. Emerges in April and feeds until June on ''Rumex'' and ''Solidago''. Pupa rounded, smooth, similar to a small bean, brownish, with dark markings; on the thorax a dark dorsal longitudinal stripe, which is continued on the abdomen by a row of impressed dots. The butterflies are on the wing from the end of June into August on meadows, clearings in woods, mountain-sides and flowery slopes; they are abundant almost everywhere in the area aru:! ascend in the high mountains above 10,000 ft.Seitz, A. ed. Band 1: Abt. 1, ''Die Großschmetterlinge des palaearktischen Faunengebietes, Die palaearktischen Tagfalter'', 1909, 379 Seiten, mit 89 kolorierten Tafeln (3470 Figuren)


Gallery

File:Lycaenidae - Lycaena virgaureae-female.JPG, ''Lycaena virgaureae'', female - lateral view File:Lycaena virgaureae female.JPG, ''Lycaena virgaureae'', female - dorsal view File:Lycaenidae - Lycaena virgaureae - male-2.JPG, ''Lycaena virgaureae'', male - lateral view File:Lycaenidae - Lycaena virgaureae - male.JPG, ''Lycaena virgaureae'', male - dorsal view File:The Macrolepidoptera of the world (Taf. 76) (8145299572).jpg , Seitz


References


External links


www.eurobutterflies.com


* ttp://www.lepiforum.de/cgi-bin/lepiwiki.pl?Lycaena_Virgaureae www.lepiforum.de: Fotos und Taxonomie
www.schmetterling-raupe.de




{{Taxonbar, from=Q520934 Lycaena Butterflies of Asia Butterflies of Europe Insects of Mongolia Butterflies described in 1758 Taxa named by Carl Linnaeus