Liana Petruțiu
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A liana is a long- stemmed woody
vine A vine is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas, or runners. The word ''vine'' can also refer to such stems or runners themselves, for instance, when used in wicker work.Jackson; Benjamin; Da ...
that is rooted in the soil at ground level and uses trees, as well as other means of vertical support, to climb up to the
canopy Canopy may refer to: Plants * Canopy (biology), aboveground portion of plant community or crop (including forests) * Canopy (grape), aboveground portion of grapes Religion and ceremonies * Baldachin or canopy of state, typically placed over an a ...
in search of direct sunlight. The word ''liana'' does not refer to a
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
grouping, but rather a habit of plant growth—much like ''
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
'' or ''
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
''. It comes from standard French , itself from an Antilles French dialect word meaning to
sheave Sheave without a rope A pulley is a wheel on an axle or shaft enabling a taut cable or belt passing over the wheel to move and change direction, or transfer power between itself and a shaft. A pulley may have a groove or grooves between flan ...
.


Ecology

Lianas are characteristic of
tropical moist broadleaf forests Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF). Description TSMF is generally found in large ...
(especially seasonal forests), but may be found in temperate rainforests and temperate deciduous forests. There are also
temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...
lianas, for example the members of the ''
Clematis ''Clematis'' is a genus of about 380 species within the buttercup family, Ranunculaceae. Their garden hybrids and cultivars have been popular among gardeners, beginning with ''Clematis'' 'Jackmanii', a garden staple since 1862; more cultivars ...
'' or ''
Vitis ''Vitis'' (grapevine) is a genus of 81 accepted species of vining plants in the flowering plant family Vitaceae. The genus consists of species predominantly from the Northern Hemisphere. It is economically important as the source of grapes, bot ...
'' (wild grape) genera. Lianas can form bridges in the forest canopy, providing
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
animals—including ants and many other invertebrates, lizards, rodents, sloths, monkeys, and lemurs—with paths through the forest. For example, in the Eastern
tropical forest Tropical forests are forested ecoregions with tropical climates – that is, land areas approximately bounded by the Tropic of Cancer, tropics of Cancer and Tropic of Capricorn, Capricorn, but possibly affected by other factors such as prevailing ...
s of
Madagascar Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
many
lemur Lemurs ( ; from Latin ) are Strepsirrhini, wet-nosed primates of the Superfamily (biology), superfamily Lemuroidea ( ), divided into 8 Family (biology), families and consisting of 15 genera and around 100 existing species. They are Endemism, ...
s achieve higher mobility from the web of lianas draped among the vertical tree species. Many lemurs prefer trees with lianas because of their roots. Lianas do not derive nutrients directly from trees, but live on and indirectly derive nutrients at the expense of trees. Specifically, they greatly reduce tree growth and tree reproduction, greatly increase tree mortality, prevent tree seedlings from establishing, alter the course of regeneration in forests, and ultimately decrease tree population growth rates. For example, forests without lianas grow 150% more fruit, and trees with lianas have twice the probability of dying. Lianas are uniquely adapted to living in forests as they use host trees, for stability, to reach to top of the canopy. Lianas directly damage their hosts by mechanical abrasion and strangulation, render hosts more susceptible to ice and wind damage, and increase the probability that the host tree falls. Lianas also provide support for weaker trees when strong winds blow by laterally anchoring them to stronger trees. However, this anchoring can also be destructive because when one tree falls, the connections made by the lianas can cause many other trees to fall. Because of these negative effects, trees that remain free of lianas are at an advantage; some species have evolved characteristics which help them avoid or shed lianas. Some lianas attain great length, such as ''
Bauhinia ''Bauhinia'' () is a large genus of flowering plants in the subfamily Cercidoideae and tribe Bauhinieae, in the large flowering plant family Fabaceae, with a pantropical distribution. The genus was named after the Bauhin brothers Gaspard and ...
'' sp. in Surinam which has grown as long as 600 meters (2000'). Hawkins has accepted a length of 1.5 km (1 mile) for an ''
Entada phaseoloides ''Entada phaseoloides'', commonly known in English as the matchbox bean or St. Thomas' bean, is a large twining vine or liana in the pea and bean family Fabaceae, native to a broad area of Asia-Pacific, from China to northern Australia and the so ...
''. The longest monocot liana is ''
Calamus manan ''Calamus manan'', the manau rattan or rotan manau, is a species of flowering plant in the palm family Arecaceae, native to Thailand, Peninsular Malaysia, Sumatra, and Borneo. A vine, its single stem is widely harvested from the wild for cane fur ...
'' (or ''Calamus ornatus'') at 240 meters (787'). Dr. Francis E. Putz states that lianas (species not indicated) have weighed "hundreds of tons" and been a half mile (0.8 km) in length. One way of distinguishing lianas from trees and shrubs is their
stiffness Stiffness is the extent to which an object resists deformation in response to an applied force. The complementary concept is flexibility or pliability: the more flexible an object is, the less stiff it is. Calculations The stiffness, k, of a ...
, specifically, the
Young's modulus Young's modulus (or the Young modulus) is a mechanical property of solid materials that measures the tensile or compressive stiffness when the force is applied lengthwise. It is the modulus of elasticity for tension or axial compression. Youn ...
of various parts of the stem. Trees and shrubs have young twigs and smaller branches that are quite flexible and older growth such as trunks and large branches that are stiffer. A liana often has stiff young growths and older, more flexible growth at the base of the stem.


Examples

Some families and genera containing liana species include:


References


External links


Lianas and Climbing Plants of the Neotropics

Lianas and Climbing Plants of the Neotropics: Family Treatments
* 'Vines and Lianas' by Rhett Butler, at http://rainforests.mongabay.com/0406.htm * {{botany Plant morphology Biology terminology Plant life-forms Plants by habit