''Leyesaurus'' is an
extinct genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
massospondylid sauropodomorph
Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had l ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
known from the
San Juan Province, northwestern
Argentina
Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, t ...
.
Description

''Leyesaurus'' is known from the
holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...
PVSJ 706, a nearly complete
skull
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, t ...
with articulated
mandible
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower teeth in place. The mandible sits beneath the maxilla. It is the only movable bon ...
and some
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated s ...
l remains (
vertebral column
The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...
,
scapular
The scapular (from Latin ''scapulae'', "shoulders") is a Western Christian garment suspended from the shoulders. There are two types of scapulars, the monastic and devotional scapular, although both forms may simply be referred to as "scapular ...
and
pelvic girdle
The pelvis (plural pelves or pelvises) is the lower part of the trunk, between the abdomen and the thighs (sometimes also called pelvic region), together with its embedded skeleton (sometimes also called bony pelvis, or pelvic skeleton).
The ...
s and
hindlimb
A hindlimb or back limb is one of the paired articulated appendages (limbs) attached on the caudal ( posterior) end of a terrestrial tetrapod vertebrate's torso.http://www.merriam-webster.com/medical/hind%20limb, Merriam Webster Dictionary-Hindl ...
). The skull has a length of 18
centimeter
330px, Different lengths as in respect to the Electromagnetic spectrum, measured by the Metre and its deriveds scales. The Microwave are in-between 1 meter to 1 millimeter.
A centimetre (international spelling) or centimeter (American spellin ...
s, and ''Leyesaurus'' has been estimated to have been about in length. It was collected from the uppermost part of the
Quebrada del Barro Formation of the
Marayes-El Carrizal Basin, dating to the
Lower Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma&nbs ...
(based on the similarities between ''Leyesaurus'' and Lower Jurassic sauropodomorphs, like ''
Massospondylus
''Massospondylus'' ( ; from Greek, (massōn, "longer") and (spondylos, "vertebra")) is a genus of sauropodomorph dinosaur from the Early Jurassic. (Hettangian to Pliensbachian ages, ca. 200–183 million years ago). It was described by S ...
'' and ''
Adeopapposaurus''). ''Leyesaurus'' was found near the locality Balde de Leyes, in the
Caucete Department of San Juan Province. Within
Massospondylidae
Massospondylidae is a family of early massopod dinosaurs that existed in Asia, Africa, North America, South America and AntarcticaHellert, Spencer M. "A New Basal Sauropodomorph from The Early Jurassic Hanson Formation of Antarctica." Geologica ...
, ''Leyesaurus'' was found to be most closely related to ''
Adeopapposaurus''.
[
]
Etymology
''Leyesaurus'' was first named by Cecilia Apaldetti, Ricardo N. Martinez, Oscar A. Alcober and Diego Pol in 2011 and the type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''Leyesaurus marayensis''. The generic name honors the Leyes family, inhabitants of the small town Balde de Leyes, who discovered the holotype and notified the paleontologists of the San Juan Museum, and ''saurus'', Greek
Greek may refer to:
Greece
Anything of, from, or related to Greece, a country in Southern Europe:
*Greeks, an ethnic group.
*Greek language, a branch of the Indo-European language family.
**Proto-Greek language, the assumed last common ancestor ...
for "lizard". The specific name refers to the Marayes-El Carrizal Basin, where the holotype was discovered.[
]
Phylogeny
The following cladogram shows the position of ''Leyesaurus'' within Massopoda, according to Oliver W. M. Rauhut and colleagues, 2020:
References
Massospondylidae
Dinosaur genera
Early Jurassic dinosaurs of South America
Jurassic Argentina
Fossils of Argentina
Geology of San Juan Province, Argentina
Fossil taxa described in 2011
{{jurassic-reptile-stub