In biology, an integument is the tissue surrounding an organism's body or an organ within, such as

skin Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other animal coverings, such as the arthropod exoskeleton, have different de ...

, a husk,

shell Shell may refer to: Architecture and design * Shell (structure), a thin structure ** Concrete shell, a thin shell of concrete, usually with no interior columns or exterior buttresses ** Thin-shell structure Science Biology * Seashell, a hard o ...

, germ or rind.

Etymology

The term is derived from ''integumentum'', which is

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power of the ...

for "a covering". In a transferred, or figurative sense, it could mean a

cloak A cloak is a type of loose garment worn over clothing, mostly but not always as outerwear for outdoor wear, serving the same purpose as an overcoat, protecting the wearer from the weather. It may form part of a uniform. Cloaks have been and ...

or a

disguise A disguise can be anything which conceals or changes a person's physical appearance, including a wig, glasses, makeup, fake moustache, costume or other items. Camouflage is a type of disguise for people, animals and objects. Hats, glasses, ch ...

. In English, "integument" is a fairly modern word, its origin having been traced back to the early seventeenth century; and refers to a material or layer with which anything is enclosed, clothed, or covered in the sense of "clad" or "coated", as with a skin or husk.

Botanical usage

botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...

, the term "integument" may be used as it is in zoology, referring to the covering of an organ. When the context indicates nothing to the contrary, the word commonly refers to an envelope covering the nucellus of the

ovule In seed plants, the ovule is the structure that gives rise to and contains the female reproductive cells. It consists of three parts: the '' integument'', forming its outer layer, the ''nucellus'' (or remnant of the megasporangium), and the ...

. The integument may consist of one layer (unitegmic) or two layers (bitegmic), each of which consisting of two or more layers of cells. The integument is perforated by a pore, the micropyle, through which the pollen tube can enter. It may develop into the testa, or

seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

coat.

Zoological usage

The integument of an organ in

zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...

typically would comprise membranes of connective tissue such as those around a kidney or liver. In referring to the integument of an animal, the usual sense is its skin and its derivatives: the integumentary system, where "integumentary" is a synonym of "

cutaneous Skin is the layer of usually soft, flexible outer tissue covering the body of a vertebrate animal, with three main functions: protection, regulation, and sensation. Other animal coverings, such as the arthropod exoskeleton, have different d ...

". In arthropods, the integument, or external "skin", consists of a single layer of

epithelial Epithelium or epithelial tissue is one of the four basic types of animal tissue, along with connective tissue, muscle tissue and nervous tissue. It is a thin, continuous, protective layer of compactly packed cells with a little intercell ...

ectoderm from which arises the cuticle, an outer covering of chitin, the rigidity of which varies as per its chemical composition.

Derivative terms and sundry usages

Derivative terms include various adjectival forms such as integumentary (e.g. system), integumental (e.g. integumental glands, "peltate glands, the integument being raised like a bladder due to abundant secretion"Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928) and integumented (as opposed to bare). Other illustrative examples of usage occur in the following articles: *

Connective tissue in skeletal muscle Skeletal muscles (commonly referred to as muscles) are organs of the vertebrate muscular system and typically are attached by tendons to bones of a skeleton. The muscle cells of skeletal muscles are much longer than in the other types of muscle ...

Dorsal artery of the penis The dorsal artery of the penis is an artery on the top surface of the penis. It is a branch of the internal pudendal artery. It runs forward on the dorsum of the penis to the glans, where it divides into two branches to the glans penis and the ...

(example of integument enclosing an internal organ) *

Flensing Flensing is the removing of the blubber or outer integument of whales, separating it from the animal's meat. Processing the blubber (the subcutaneous fat) into whale oil was the key step that transformed a whale carcass into a stable, transpor ...

Flesh Flesh is any aggregation of soft tissues of an organism. Various multicellular organisms have soft tissues that may be called "flesh". In mammals, including humans, ''flesh'' encompasses muscles, fats and other loose connective tissues, but ...

(generic use of plural "integuments") *

Herzog & de Meuron Herzog & de Meuron Basel Ltd.,
" Herzog & de Meuron. Retrieved on 11 October 2012. "Herzog & de Meuron Basel Ltd. R ...

(figurative usage) * Integumentary system (senses related to vertebrate skin and medical aspects) * Integumental muscles *

Mosasaur Mosasaurs (from Latin ''Mosa'' meaning the 'Meuse', and Greek ' meaning 'lizard') comprise a group of extinct, large marine reptiles from the Late Cretaceous. Their first fossil remains were discovered in a limestone quarry at Maastricht on ...

Seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiospe ...

References

{{Authority control Animal anatomy Skin Organs (anatomy) Plant anatomy