Hart's-tongue Thyme-moss
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''Asplenium scolopendrium'', commonly known as the hart's-tongue fern, is an
evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...
fern The ferns (Polypodiopsida or Polypodiophyta) are a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. They differ from mosses by being vascular, i.e., having specialized tissue ...
in the
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Aspleniaceae The Aspleniaceae (spleenworts) are a family of ferns, included in the order Polypodiales. The composition and classification of the family have been subject to considerable changes. In particular, there is a narrow circumscription, Aspleniaceae ...
native to the Northern Hemisphere.


Description

The most striking and unusual feature of the fern is its simple, strap-shaped undivided
frond A frond is a large, divided leaf. In both common usage and botanical nomenclature, the leaves of ferns are referred to as fronds and some botanists restrict the term to this group. Other botanists allow the term frond to also apply to the lar ...
s. The supposed resemblance of the leaves to the tongue of a
hart Hart often refers to: * Hart (deer) * Hart (surname) Hart may also refer to: Organizations * Hart Racing Engines, a former Formula One engine manufacturer * Hart Skis, US ski manufacturer * Hart Stores, a Canadian chain of department store ...
(an archaic term for a male
red deer The red deer (''Cervus elaphus'') is one of the largest deer species. A male red deer is called a stag or Hart (deer), hart, and a female is called a doe or hind. The red deer inhabits most of Europe, the Caucasus Mountains region, Anatolia, Ir ...
) gave rise to the common name "hart's-tongue fern".


Taxonomy

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
first gave the hart's-tongue fern the
binomial Binomial may refer to: In mathematics *Binomial (polynomial), a polynomial with two terms *Binomial coefficient, numbers appearing in the expansions of powers of binomials *Binomial QMF, a perfect-reconstruction orthogonal wavelet decomposition * ...
''Asplenium scolopendrium'' in his ''
Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genus, genera. It is the first work to consistently apply binomial nomenclature ...
'' of 1753. The Latin
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
''scolopendrium'' is derived from the Greek ''skolopendra'', meaning a centipede or millipede; this is due to the sori pattern being reminiscent of a
myriapod Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial. Although molecular evidence and similar fossils suggests a diversifi ...
's legs. A global phylogeny of ''Asplenium'' published in 2020 divided the genus into eleven clades, which were given informal names pending further taxonomic study. ''A. scolopendrium'' belongs to the "''Phyllitis'' subclade" of the "''Phyllitis'' clade". Members of the ''Phyllitis'' clade have undivided or pinnatifid leaf blades with a thick, leathery texture, persistent scales on their stalk, and often possess
anastomosing An anastomosis (, : anastomoses) is a connection or opening between two things (especially cavities or passages) that are normally diverging or branching, such as between blood vessels, leaf veins, or streams. Such a connection may be normal (suc ...
veins. Members of the ''Phyllitis'' subclade have undivided leaves with freely branching veins and single or paired sori. They are widely distributed through the Northern Hemisphere. If defined to include the former '' A. komarovii'', ''A. scolopendrium'' makes up the former segregate genus ''Phyllitis'' and is sister to '' A. sagittatum''. Three subspecies were accepted in the most recent revision of the species: *''A. scolopendrium'' subsp. ''scolopendrium'' is native to
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
,
North Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
, the
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
, and
Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
. Populations in the eastern Mediterranean have been referred to as ''A. scolopendrium'' var. ''antri-jovis''. *''A. scolopendrium'' subsp. ''americanum'' is native to
Ontario Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...
in
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, southern
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, and
Hispaniola Hispaniola (, also ) is an island between Geography of Cuba, Cuba and Geography of Puerto Rico, Puerto Rico in the Greater Antilles of the Caribbean. Hispaniola is the most populous island in the West Indies, and the second-largest by List of C ...
. The tropical populations have been referred to as ''A. scolopendrium'' var. ''lindenii''. *''A. scolopendrium'' subsp. ''japonicum'', formerly ''A. komarovii'', is native to eastern
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
, and
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
Morphological differences between the varieties are minor, but the North American variety ''americanum'' is
tetraploid Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
, while the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
subspecies ''A. scolopendrium scolopendrium'') is
diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...
.


Distribution

''Asplenium scolopendrium'' is a common species in the Old World: *The subspecies ''scolopendrium'' occurs throughout
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
(including the
Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
and the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
). It is absent however in northeast Europe (Finland, Belarus, Latvia, Lithuania and European Russia).Christenhusz, M., Bento Elias, R., Dyer, R., Ivanenko, Y., Rouhan, G., Rumsey, F. & Väre, H. 2017. ''Asplenium scolopendrium (Europe assessment)''. ''The IUCN Red List of Threatened Species'' 2017: e.T83504456A83506952. Accessed on 19 June 2024. Beyond Europe, it is found in the
Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...
, and North Africa (in
Morocco Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
,
Algeria Algeria, officially the People's Democratic Republic of Algeria, is a country in the Maghreb region of North Africa. It is bordered to Algeria–Tunisia border, the northeast by Tunisia; to Algeria–Libya border, the east by Libya; to Alger ...
,
Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...
and
Libya Libya, officially the State of Libya, is a country in the Maghreb region of North Africa. It borders the Mediterranean Sea to the north, Egypt to Egypt–Libya border, the east, Sudan to Libya–Sudan border, the southeast, Chad to Chad–L ...
, as well as the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
). It has also been introduced to the
Falkland Islands The Falkland Islands (; ), commonly referred to as The Falklands, is an archipelago in the South Atlantic Ocean on the Patagonian Shelf. The principal islands are about east of South America's southern Patagonian coast and from Cape Dub ...
. Unlike its American counterpart, the ''scolopendrium'' of Europe is used for many horticultural uses. Specimens of this variety found in North America (such as in
New Brunswick New Brunswick is a Provinces and Territories of Canada, province of Canada, bordering Quebec to the north, Nova Scotia to the east, the Gulf of Saint Lawrence to the northeast, the Bay of Fundy to the southeast, and the U.S. state of Maine to ...
and
Ontario Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...
in Canada, and
Maryland Maryland ( ) is a U.S. state, state in the Mid-Atlantic (United States), Mid-Atlantic region of the United States. It borders the states of Virginia to its south, West Virginia to its west, Pennsylvania to its north, and Delaware to its east ...
in the United States) are considered naturalized descendants of cultivated plants. In
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, it occurs in rare, widely scattered populations located in different locales: * in the Mexican states of
Chiapas Chiapas, officially the Free and Sovereign State of Chiapas, is one of the states that make up the Political divisions of Mexico, 32 federal entities of Mexico. It comprises Municipalities of Chiapas, 124 municipalities and its capital and large ...
,
Coahuila Coahuila, formally Coahuila de Zaragoza, officially the Free and Sovereign State of Coahuila de Zaragoza, is one of the 31 states of Mexico. The largest city and State Capital is the city of Saltillo; the second largest is Torreón and the thi ...
,
Nuevo León Nuevo León, officially the Free and Sovereign State of Nuevo León, is a Administrative divisions of Mexico, state in northeastern Mexico. The state borders the Mexican states of Tamaulipas, Coahuila, Zacatecas, and San Luis Potosí, San Luis ...
, and
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
, as well as the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
island of
Hispaniola Hispaniola (, also ) is an island between Geography of Cuba, Cuba and Geography of Puerto Rico, Puerto Rico in the Greater Antilles of the Caribbean. Hispaniola is the most populous island in the West Indies, and the second-largest by List of C ...
(in
Haiti Haiti, officially the Republic of Haiti, is a country on the island of Hispaniola in the Caribbean Sea, east of Cuba and Jamaica, and south of the Bahamas. It occupies the western three-eighths of the island, which it shares with the Dominican ...
). * along the
Onondaga Limestone The Onondaga Limestone is a group of hard limestones and dolomites of Devonian age that forms geographic features in some areas in which it outcrops; in others, especially its Southern Ontario portion, the formation can be less prominent as a lo ...
and
Niagara Escarpment The Niagara Escarpment is an approximately discontinuous, arc-shaped but generally northward-facing escarpment, or cuesta, in Canada and the United States. The escarpment begins south of Lake Ontario and circumscribes the top of the Great Lake ...
geological formations in
Central New York The central region of New York state includes: * Auburn in Cayuga County * Cortland in Cortland County * Oneida in Madison County * Syracuse, the largest city of Central New York, in Onondaga County * Fulton and Oswego in Oswego County ...
(present in 2 counties),
southern Ontario Southern Ontario is a Region, primary region of the Canadian Provinces and territories of Canada, province of Ontario. It is the most densely populated and southernmost region in Canada, with approximately 13.5 million people, approximately 36% o ...
(present in 7 counties), and the eastern
Upper Peninsula The Upper Peninsula of Michigan—also known as Upper Michigan or colloquially the U.P. or Yoop—is the northern and more elevated of the two major landmasses that make up the U.S. state of Michigan; it is separated from the Lower Peninsula b ...
in
Michigan Michigan ( ) is a peninsular U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, Upper Midwestern United States. It shares water and land boundaries with Minnesota to the northwest, Wisconsin to the west, ...
(present in 2 counties). Exceptions are
disjunct population In biology, a taxon with a disjunct distribution is one that has two or more groups that are related but considerably separated from each other geographically. The causes are varied and might demonstrate either the expansion or contraction of a s ...
s that exist in
Alabama Alabama ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South, Deep Southern regions of the United States. It borders Tennessee to the north, Georgia (U.S. state), Georgia to the east, Florida and the Gu ...
(in Fern Cave National Wildlife Refuge, a wildlife refuge centered around an off-limits cave in
Jackson County, Alabama Jackson County is the northeasternmost County (United States), county in the U.S. state of Alabama. As of the 2020 United States census, 2020 census, the population was 52,579. The county seat is Scottsboro, Alabama, Scottsboro. The county was ...
, where it has declined heavily due to illegal plant collecting, and an undisclosed pit in Morgan County that is also off-limits and protected), and
Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...
(in just a single county); these southern populations are at dire risk of
extirpation Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with extinction, global extinctions. Local extinctions ...
. An introduced population descended from New York plants is found in
New Jersey New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
; it is a remnant of a 1936 effort to practice
ex-situ conservation Svalbard Global Seed Bank, an ' conservation ''Ex situ'' conservation () is the process of protecting an endangered species, variety, or breed of plant or animal outside its natural habitat. For example, by removing part of the population from ...
of populations in New York. * In 2020, a new population of hart's-tongue ferns was discovered inside a cave with basaltic lava flows in
El Malpais National Monument El Malpais National Monument is a National Monument located in western New Mexico, in the Southwestern United States. The name El Malpais is from the Spanish term '' Malpaís'', meaning ''badlands'', due to the extremely barren and dramatic volc ...
,
Cibola County, New Mexico Cibola County is a county in the U.S. state of New Mexico. As of the 2020 census, the population was 27,172. Its county seat is Grants. It is New Mexico's youngest county, and the third youngest county in the United States, created on June ...
; this represents the first confirmed population of the species in North America west of the Mississippi. Genetic analyses and surveys are currently being performed to determine the population's variation and overall health. In East Asia, ''A. scolopendrium'' subsp. ''japonicum'' is distributed in the Russian Far East, north-eastern China and the Korean Peninsula, however it is considered as being generally rare on the mainland. In contrast, it is relatively abundant on many islands including the Japanese Archipelago. The unique dispersal of ''Asplenium scolopendrium'' has caught the attention of international botanists. In fact, the very existence of such varieties beg that "...these populations arose following colonization events involving a single spore".


Habitat

The plants grow on neutral, calcium-rich, and/or lime-rich substrates under deciduous hardwood canopies (usually
sugar maple ''Acer saccharum'', the sugar maple, is a species of flowering plant in the soapberry and lychee family Sapindaceae. It is native to the hardwood forests of eastern Canada and the eastern United States. Sugar maple is best known for being the ...
s in eastern North America), including moist soil and damp crevices in old walls; they are found most commonly in shaded areas. Plants in full sun are usually stunted and yellowish in colour, while those in full shade are dark green and healthy. The disjunct populations of the North American variation in the southeastern US are found exclusively in sinkhole pits or limestone caves. These populations may be relics of cooler
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
climates.


Conservation

The most recent
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the stat ...
(IUCN) assessment for
The IUCN Red List of Threatened Species The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological spe ...
was conducted in 2016, with the conservation status of ''Asplenium scolopendrium'' being listed as being a species of Least Concern (LC). Despite the positive status, this assessment is chiefly limited to the conservation situation of ''A. scolopendrium'' in Europe, as opposed to other parts of its natural range.


Europe

''Asplenium scolopendrium'' is found across the vast majority of European countries, in a wide variety of natural settings, across a multitude of habitats. It is also able to colonise manmade environments including growing in walls and brickwork crevices, particularly in limestone mortar. Furthermore, it is considered to be a gregarious species – it is able to spread both sexually and asexually, frequently forming large clumps and present in high densities – in some areas it is considered very common. Across Europe as a whole, the IUCN assessed no immediate threats to this species in natural settings, and predicted that the species may benefit from milder wetter winters predicted under various climate change scenarios. In the urban environment, where ''A. scolopendrium'' frequently grows on vertical faces and in crevices of lime mortared brick walls, the species may be threatened by wall renovation projects. Although the fern is often cultivated as a garden plant or used in landscape planting, commercially sold plants are sourced from horticultural stock. ''Asplenium scolopendrium'' is not a specifically protected species in the majority of European states, however in part due to its wide distribution, ''A. scolopendrium'' inhabits a large number of legally protected areas across the continent. Despite this, it is considered to be declining in numbers or a species at risk in certain parts of its range. ''Asplenium scolopendrium'' was first listed as "Vulnerable" in the National Red Lists for
Albania Albania ( ; or ), officially the Republic of Albania (), is a country in Southeast Europe. It is located in the Balkans, on the Adriatic Sea, Adriatic and Ionian Seas within the Mediterranean Sea, and shares land borders with Montenegro to ...
in 2014 and
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. The remote Arctic island of Jan Mayen and the archipelago of Svalbard also form part of the Kingdom of ...
in 2010 (under Criterion D1, that is, having a very small population of under 1000 individuals). It is considered "critically threatened and rare" in the
Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...
's 2012 plant Red List; and "Endangered" in
Sweden Sweden, formally the Kingdom of Sweden, is a Nordic countries, Nordic country located on the Scandinavian Peninsula in Northern Europe. It borders Norway to the west and north, and Finland to the east. At , Sweden is the largest Nordic count ...
's 2010 Red List.


North America


United States

In the United States, ''A. scolopendrium'' var. ''americanum'' was declared endangered by the U.S. Fish and Wildlife Service in 1989. The reasons for its rarity are currently being researched, with
reintroduction Species reintroduction is the deliberate release of a species into the wild, from captivity or other areas where the organism is capable of survival. The goal of species reintroduction is to establish a healthy, genetically diverse, self-sustain ...
programs in New York and elsewhere also in development.


Canada

Ontario, Canada has the highest population numbers of ''A. scolopendrium'' var. ''americanum'' of any region in the variation's distribution, with around 80% of all subpopulations and around 94% of all individuals. The fern was reported at more than 100 sites across the province, with around 75 still believed to be existing. Despite this, ''A. scolopendrium'' var. ''americanum'' was listed as a species of Special Concern under the Committee on the Status of Species at Risk in Ontario in May 2017, due to its extremely specific habitat requirements, relatively small distribution, and some subpopulations consisting of too little individuals.


Uses


Cultivation

''Asplenium scolopendrium'' is often grown as an
ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
, with several
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s selected with varying frond form, including with frilled frond margins, forked fronds and cristate forms. The species has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
, as has the cultivar 'Angustatum'. The American variety is reputed to be difficult to cultivate (making conservation efforts for it even more troublesome); due to this, most, if not all, cultivated individuals are derived from the Old World subspecies.


Herbal medicine

This fern was used in the 1800s as a
medicinal plant Medicinal plants, also called medicinal herbs, have been discovered and used in traditional medicine practices since prehistoric times. Plants synthesize hundreds of chemical compounds for various functions, including Plant defense against h ...
in folk medicine as a
spleen The spleen (, from Ancient Greek '' σπλήν'', splḗn) is an organ (biology), organ found in almost all vertebrates. Similar in structure to a large lymph node, it acts primarily as a blood filter. The spleen plays important roles in reg ...
tonic (hence an archaic name for the genus, "spleenworts") and for other uses.


Other uses

A manuscript written by Katharine Palmer in the 18th century contains a recipe for hart's-tongue ale.


Gallery

Phyllitis scolopendrium habitat.jpg, Habitat Phyllitis scolopendrium3.jpg, Spiers harts tongue fern.JPG, Asplenium scolopendrium americanum.jpg,


References

* *


Further reading

* Hyde, H. A., Wade, A. E., & Harrison, S. G. (1978). ''Welsh Ferns''. National Museum of Wales. . * A popular article on hart's tongue fern that includes several references and a discussion of cultivation possibilities for the European and American varieties. The article strongly discourages collection and or cultivation of the North American variety.


External links


Commentary and video on Hart's Tongue ferns at Lynn Glen, Dalry, Scotland
{{Authority control scolopendrium Garden plants of North America Medicinal plants of North America Plants described in 1753 Ferns of Europe Ferns of Asia Ferns of the Americas Taxa named by Carl Linnaeus