Haplogroup M is a
human mitochondrial DNA (mtDNA)
haplogroup. An enormous haplogroup spanning all the continents, the macro-haplogroup M, like its sibling the
macro-haplogroup N, is a descendant of the
haplogroup L3.
All mtDNA haplogroups considered native outside of Africa are descendants of either haplogroup M or its sibling haplogroup N.
Haplogroup M is relatively young, having a younger
most recent common ancestor date than some subclades of haplogroup N such as
haplogroup R.
Origins
There is a debate concerning the geographical origins of Haplogroup M and its sibling haplogroup N. Both lineages are thought to have been the main surviving lineages involved in the
out of Africa migration (or migrations) because all indigenous lineages found outside Africa belong to haplogroup M or haplogroup N. Scientists are unsure whether the mutations that define haplogroups M and N occurred in Africa before the exit from Africa or in Asia after the exit from Africa. Determining the origins of haplogroup M is further complicated by an early back-migration (from Asia to Africa) of bearers of M1.
Its date of origin in absolute terms is only known with great uncertainty, as reconstruction has yielded different (but overlapping) ranges for the age of M in South Asia and
East Asia
East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of Economy of China, China, Economy of Ja ...
. The same authors give an estimate for t of L3 as , later (2011) narrowed to the somewhat younger .
Thus, haplogroup M would have emerged around 10,000 or at most 20,000 years after L3, around or somewhat after the
recent out-of-Africa migration event.
Haplogroup M1
Much discussion concerning the origins of haplogroup M has been related to its subclade haplogroup M1, which is the only variant of macro-haplogroup M found in Africa.
Two possibilities were being considered as potential explanations for the presence of M1 in Africa:
# M was present in the ancient population which later gave rise to both M1 in Africa, and M more generally found in
Eurasia
Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
.
# The presence of M1 in Africa is the result of a back-migration from Asia which occurred sometime after the Out of Africa migration.
Haplogroup M23
In 2009, two independent publications reported a rare, deep-rooted subclade of haplogroup M, referred to as M23, that is present in
Madagascar
Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
.
The origin of M23 is still not clearly understood, a west Eurasian origin had been suggested before. Haplogroup M23 is ancient and found throughout Madagascar. It is suggested to be an Asian component with significant African component. M23 was found only in one individual in Dubai
Asian origin hypothesis
According to this theory, anatomically modern humans carrying ancestral haplogroup L3 lineages were involved in the Out of Africa migration from East Africa into Asia. Somewhere in Asia, the ancestral L3 lineages gave rise to haplogroups M and N. The ancestral L3 lineages were then lost by
genetic drift
Genetic drift, also known as random genetic drift, allelic drift or the Wright effect, is the change in the Allele frequency, frequency of an existing gene variant (allele) in a population due to random chance.
Genetic drift may cause gene va ...
as they are infrequent outside Africa. The hypothesis that Asia is the origin of macrohaplogroup M is supported by the following:
#The highest frequencies worldwide of macrohaplogroup M are observed in Asia, specifically in the Indian subcontinent, China, Japan and Korea where frequencies range from 60 to 80%. The total frequency of M subclades is even higher in some populations of Siberia or the Americas, but these small populations tend to exhibit strong genetic drift effects, and often their geographical neighbors exhibit very different frequencies.
#Deep time depth >50,000 years of western, central, southern and eastern Indian haplogroups M2, M38, M54, M58, M33, M6, M61, M62 and the distribution of macrohaplogroup M, do not rule out the possibility of macrohaplogroup M arising in Indian population.
#With the exception of the African specific M1, India has several M lineages that emerged directly from the root of haplogroup M.
#Only two subclades of haplogroup M, M1 and M23, are found in Africa, whereas numerous subclades are found outside Africa
(with some discussion possible only about sub-clade M1, concerning which see below).
#Specifically concerning M1
::*Haplogroup M1 has a restricted geographic distribution in Africa, being found mainly in North Africans and East Africa at low or moderate frequencies. If M had originated in Africa around, or before, the Out of Africa migration, it would be expected to have a more widespread distribution
::*According to Gonzalez et al. 2007, M1 appears to have expanded relatively recently. In this study M1 had a younger coalescence age than the Asian-exclusive M lineages.
::*The geographic distribution of M1 in Africa is predominantly North African/supra-equatorial
and is largely confined to
Afro-Asiatic
The Afroasiatic languages (also known as Afro-Asiatic, Afrasian, Hamito-Semitic, or Semito-Hamitic) are a language family (or "phylum") of about 400 languages spoken predominantly in West Asia, North Africa, the Horn of Africa, and parts of th ...
speakers,
which is inconsistent with the Sub-Saharan distribution of sub-clades of haplogroups
L3 and
L2 that have similar time depths.
::*One of the basal lineages of M1 lineages has been found in
Northwest Africa and in the
Near East
The Near East () is a transcontinental region around the Eastern Mediterranean encompassing the historical Fertile Crescent, the Levant, Anatolia, Egypt, Mesopotamia, and coastal areas of the Arabian Peninsula. The term was invented in the 20th ...
but is absent in East Africa.
::*M1 is not restricted to Africa. It is relatively common in the Mediterranean, peaking in
Iberia
The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, compri ...
. M1 also enjoys a well-established presence in the Middle East, from the south of the Arabian Peninsula to
Anatolia
Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...
and from the
Levant
The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
to
Iran
Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
. In addition, M1 haplotypes have occasionally been observed in the
Caucasus
The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...
and the Trans Caucasus, and without any accompanying L lineages.
M1 has also been detected in
Central Asia
Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...
, seemingly reaching as far as
Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
.
::*The fact that the M1 sub-clade of macrohaplogroup M has a coalescence age which overlaps with that of
haplogroup U6 (a Eurasian haplogroup whose presence in Africa is due to a back-migration from West Asia) and the distribution of U6 in Africa is also restricted to the same North African and Horn African populations as M1 supports the scenario that M1 and U6 were part of the same population expansion from Asia to Africa.
[
::*The timing of the proposed migration of M1 and U6-carrying peoples from West Asia to Africa (between 40,000 and 45,000 ybp) is also supported by the fact that it coincides with changes in climatic conditions that reduced the desert areas of North Africa, thereby rendering the region more accessible to entry from the Levant. This climatic change also temporally overlaps with the peopling of Europe by populations bearing haplogroup U5, the European sister clade of haplogroup U6.][
]
African origin hypothesis
According to this theory, haplogroups M and N arose from L3 in an East African population ancestral to eurasians that had been isolated from other African populations before the OOA event. Members of this population were involved in the out Africa migration and may have only carried M and N lineages. With the possible exception of haplogroup M1, all other M and N clades in Africa were lost due to admixture with other African populations and genetic drift
Genetic drift, also known as random genetic drift, allelic drift or the Wright effect, is the change in the Allele frequency, frequency of an existing gene variant (allele) in a population due to random chance.
Genetic drift may cause gene va ...
.
The African origin of Haplogroup M is supported by the following arguments and evidence.
#L3, the parent clade of haplogroup M, is found throughout Africa, but is rare outside Africa. According to Toomas Kivisild (2003), "the lack of L3 lineages other than M and N in India and among non-African mitochondria in general suggests that the earliest migration(s) of modern humans already carried these two mtDNA ancestors, via a departure route over the Horn of Africa."[
#Specifically concerning at least M1a:
::This study provides evidence that M1, or its ancestor, had an Asiatic origin. The earliest M1 expansion into Africa occurred in northwestern instead of northeastern areas; this early spread reached the Iberian Peninsula even affecting the Basques. The majority of the M1a lineages found outside and inside Africa had a more recent eastern Africa origin. Both western and eastern M1 lineages participated in the Neolithic colonization of the Sahara. The striking parallelism between subclade ages and geographic distribution of M1 and its North African U6 counterpart strongly reinforces this scenario. Finally, a relevant fraction of M1a lineages present today in the European Continent and nearby islands possibly had a Jewish instead of the commonly proposed Amazigh maternal ascendance.]
Dispersal
A number of studies have proposed that the ancestors of modern haplogroup M dispersed from Africa through the southern route across the Horn of Africa along the coastal regions of Asia onwards to New Guinea and Australia. These studies suggested that the migrations of haplogroups M and N occurred separately with haplogroup N heading northwards from East Africa to the Levant
The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...
. However, the results of numerous recent studies indicate that there was only one migration out of Africa and that haplogroups M and N were part of the same migration. This is based on the analysis of a number of relict populations along the proposed beachcombing route from Africa to Australia, all of which possessed both haplogroups N and M.
A 2008 study by Abu-Amero et al., suggests that the Arabian Peninsula
The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world.
Geographically, the ...
may have been the main route out of Africa. However, as the region lacks of autochthonous clades of haplogroups M and N the authors suggest that the area has been a more recent receptor of human migrations than an ancient demographic expansion center along the southern coastal route as proposed under the single migration Out-of-Africa scenario of the African origin hypothesis.
Distribution
M is the most common mtDNA haplogroup in Asia, super-haplogroup M is distributed all over Asia
Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, where it represents 60% of all maternal lineages.
All Andamanese belong to Haplogroup M. It peaks in the Malaysian aboriginal Negrito tribes at almost 100% but with mtDNA M21a representing Semang; 84% in Mendriq people, Batek people 48%, (almost all belong to the specific Malaysian Negrito haplogroup M21a, this subclade also found in the Orang Asli
The Orang Asli are a Homogeneity and heterogeneity, heterogeneous Indigenous peoples, indigenous population forming a national minority in Malaysia. They are the oldest inhabitants of Peninsular Malaysia.
As of 2017, the Orang Asli accounted f ...
21%, Thais 7.8% and Malay 4.6%) It also peaks very high in Japan and Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, where it represents on average about 70% of the maternal lineages (160/216 = 74% Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, 205/282 = 73% Tōkai, 231/326 = 71% Okinawa, 148/211 = 70% Japanese, 50/72 = 69% Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, 150/217 = 69% Hokkaidō, 24/35 = 69% Zhongdian Tibetan, 175/256 = 68% northern Kyūshū, 38/56 = 68% Qinghai
Qinghai is an inland Provinces of China, province in Northwestern China. It is the largest provinces of China, province of China (excluding autonomous regions) by area and has the third smallest population. Its capital and largest city is Xin ...
Tibetan, 16/24 = 67% Diqing Tibetan, 66/100 = 66% Miyazaki, 33/51 = 65% Ainu, 214/336 = 64% Tōhoku, 75/118 = 64% Tokyo
Tokyo, officially the Tokyo Metropolis, is the capital of Japan, capital and List of cities in Japan, most populous city in Japan. With a population of over 14 million in the city proper in 2023, it is List of largest cities, one of the most ...
(JPT)) and is ubiquitous in India and South Korea
South Korea, officially the Republic of Korea (ROK), is a country in East Asia. It constitutes the southern half of the Korea, Korean Peninsula and borders North Korea along the Korean Demilitarized Zone, with the Yellow Sea to the west and t ...
, where it has approximately 60% frequency. Among Chinese people
The Chinese people, or simply Chinese, are people or ethnic groups identified with Greater China, China, usually through ethnicity, nationality, citizenship, or other affiliation.
Chinese people are known as Zhongguoren () or as Huaren () by ...
both inside and outside of China, haplogroup M accounts for approximately 50% of all mtDNA on average, but the frequency varies from approximately 40% in Hans from Hunan
Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
and Fujian
Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
in southern China to approximately 60% in Shenyang
Shenyang,; ; Mandarin pronunciation: ; formerly known as Fengtian formerly known by its Manchu language, Manchu name Mukden, is a sub-provincial city in China and the list of capitals in China#Province capitals, provincial capital of Liaonin ...
, Liaoning
)
, image_skyline =
, image_alt =
, image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong
, image_map = Liaoning in China (+all claims hatched).svg
, ...
in northeastern China.
Haplogroup M accounts for approximately 42% of all mtDNA in Filipinos
Filipinos () are citizens or people identified with the country of the Philippines. Filipinos come from various Austronesian peoples, all typically speaking Filipino language, Filipino, Philippine English, English, or other Philippine language ...
, among whom it is represented mainly by M7c3c and E. In Vietnam, haplogroup M has been found in 37% (52/139) to 48% (20/42) of samples of Vietnamese and in 32% (54/168) of a sample of Chams from Bình Thuận Province. Haplogroup M accounts for 43% (92/214) of all mtDNA in a sample of Laotians, with its subclade M7 (M7b, M7c, and M7e) alone accounting for a full third of all haplogroup M, or 14.5% (31/214) of the total sample.
In Oceania
Oceania ( , ) is a region, geographical region including Australasia, Melanesia, Micronesia, and Polynesia. Outside of the English-speaking world, Oceania is generally considered a continent, while Mainland Australia is regarded as its co ...
, A 2008 study found Haplogroup M in 42% (60/144) of a pool of samples from nine language groups in the Admiralty Islands of Papua New Guinea. M has been found in 35% (17/48) of a sample of Papua New Guinea highlanders from the Bundi area and in 28% (9/32) of a sample of Aboriginal Australians from Kalumburu in northwestern Australia. In a study published in 2015, Haplogroup M was found in 21% (18/86) of a sample of Fijians
Fijians () are a nation and ethnic group Indigenous peoples of Oceania, native to Fiji, who speak Fijian language, Fijian and English language, English and share a common history and culture.
Fijians, or ''iTaukei'', are the major indigenous ...
, but it was not observed in a sample of 21 Rotumans.
Haplogroup M is also relatively common in Northeast Africa, occurring especially among Somalis
The Somali people (, Wadaad's writing, Wadaad: , Arabic: ) are a Cushitic peoples, Cushitic ethnic group and nation native to the Somali Peninsula. who share a common ancestry, culture and history.
The Lowland East Cushitic languages, East ...
, Libya
Libya, officially the State of Libya, is a country in the Maghreb region of North Africa. It borders the Mediterranean Sea to the north, Egypt to Egypt–Libya border, the east, Sudan to Libya–Sudan border, the southeast, Chad to Chad–L ...
ns and Oromos at frequencies over 20%. Toward the northwest, the lineage is found at comparable frequencies among the Tuareg in Mali and Burkina Faso; particularly the M1a2 subclade (18.42%).
Among the descendant lineages of haplogroup M are C, D, E, G, Q, and Z. Z and G are found in North Eurasian populations, C and D exists among North Eurasian and Native American populations, E is observed in Southeast Asian populations, and Q is common among Melanesian populations. The lineages M2, M3, M4, M5, M6, M18 and M25 are exclusive to South Asia, with M2 reported to be the oldest lineage on the Indian sub-continent with an age estimation of 60,000—75,000 years, and with M5 reported to be the most prevalent in historically Turco-Persian enclaves.
In 2013, four ancient specimens dated to around 2,500 BC-500 AD, which were excavated from the Tell Ashara ( Terqa) and Tell Masaikh (Kar-Assurnasirpal) archaeological sites in the Euphrates Valley, were found to belong to mtDNA haplotypes associated with the M4b1, M49 and/or M61 haplogroups. Since these clades are not found among the current inhabitants of the area, they are believed to have been brought at a more remote period from east of Mesopotamia
Mesopotamia is a historical region of West Asia situated within the Tigris–Euphrates river system, in the northern part of the Fertile Crescent. Today, Mesopotamia is known as present-day Iraq and forms the eastern geographic boundary of ...
; possibly by either merchants or the founders of the ancient Terqa population.
In 2016, three Late Pleistocene European hunter-gatherers were also found to carry M lineages. Two of the specimens were from the Goyet archaeological site in Belgium and were dated to 34,000 and 35,000 years ago, respectively. The other ancient individual hailed from the La Rochette site in France, and was dated to 28,000 years ago.
Ancient DNA analysis of Iberomaurusian skeletal remains at the Taforalt
Taforalt, or Grotte des Pigeons, is a cave in the province of Berkane, Aït Iznasen region, Morocco, possibly the oldest cemetery in North Africa. It contained at least 34 Iberomaurusian adolescent and adult human skeletons, as well as young ...
site in Morocco, which have been dated to between 15,100 and 13,900 ybp, observed the M1b subclade in one of the fossils (1/7; ~14%). Ancient individuals belonging to the Late Iron Age settlement of Çemialo Sırtı in Batman
Batman is a superhero who appears in American comic books published by DC Comics. Batman was created by the artist Bob Kane and writer Bill Finger, and debuted in Detective Comics 27, the 27th issue of the comic book ''Detective Comics'' on M ...
, southeast Turkey were found to carry haplogroup M; specifically the M1a1 subclade (1/12; ~8.3%). Haplogroup M was also detected in ancient specimens from Southeast Anatolia (0.4%). Additionally, M1 has been observed among ancient Egypt
Ancient Egypt () was a cradle of civilization concentrated along the lower reaches of the Nile River in Northeast Africa. It emerged from prehistoric Egypt around 3150BC (according to conventional Egyptian chronology), when Upper and Lower E ...
ian mummies excavated at the Abusir el-Meleq archaeological site in Middle Egypt, which date from the Pre- Ptolemaic/late New Kingdom and Roman periods. Fossils at the Early Neolithic site of Ifri n'Amr or Moussa in Morocco
Morocco, officially the Kingdom of Morocco, is a country in the Maghreb region of North Africa. It has coastlines on the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to Algeria–Morocc ...
, which have been dated to around 5,000 BCE, have also been found to carry the M1b subclade. These ancient individuals bore an autochthonous Maghrebi genomic component that peaks among modern Berbers
Berbers, or the Berber peoples, also known as Amazigh or Imazighen, are a diverse grouping of distinct ethnic groups indigenous to North Africa who predate the arrival of Arab migrations to the Maghreb, Arabs in the Maghreb. Their main connec ...
, indicating that they were ancestral to populations in the area. The ancient Egyptian aristocrats Nakht-Ankh and Khnum-Nakht were also found to belong to the M1a1 subclade. The half-brothers lived during the 12th Dynasty, with their tomb
A tomb ( ''tumbos'') or sepulchre () is a repository for the remains of the dead. It is generally any structurally enclosed interment space or burial chamber, of varying sizes. Placing a corpse into a tomb can be called '' immurement'', alth ...
located at the Deir Rifeh cemetery in Middle Egypt.
Subgroups distribution
Source:
* M
** M1'20'51
*** Haplogroup M
– found in the Nile Valley, Horn of Africa, North Africa
North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...
, Sahara
The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...
, Mediterranean
The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
, and Middle East
The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq.
The term came into widespread usage by the United Kingdom and western Eur ...
**** M1a
***** M1a1
****** M1a1a
******* M1a1a1
****** M1a1b
******* M1a1b1
******** M1a1b1a
******** M1a1b1b
********* M1a1b1b1
******** M1a1b1c
******* M1a1b2
****** M1a1-T16093C
******* M1a1c
******* M1a1d
****** M1a1e
******* M1a1e1
******* M1a1e2
****** M1a1f
****** M1a1g
****** M1a1h
****** M1a1i
***** M1a2
****** M1a2a
****** M1a2b
***** M1a3
****** M1a3a
****** M1a3b
******* M1a3b1
******* M1a3b2
***** M1a4
****** M1a4a
***** M1a5
***** M1a6
***** M1a7
***** M1a8
****** M1a8a
**** M1b
***** M1b1
****** M1b1a
****** M1b1b
***** M1b2
****** M1b2a
****** M1b2b
****** M1b2c
*** M20 – in China, Borneo (Bidayuh), Thailand, Laos, Indonesia, Vietnam ( Jarai, Lahu), Saudi Arabia
**** M20a - Myanmar, Thailand ( Shan from Mae Hong Son Province, Tai Yuan), China, Blang, Saudi Arabia,
***** M20a1 - Thailand ( Lawa from southeastern Mae Hong Son Province[YFull MTree 1.01.14609](_blank)
as of 21 August 2019), Myanmar
***** M20a2 - Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
( Phutai from Kalasin Province, Nyaw from Nakhon Phanom Province, Bru from Sakon Nakhon Province, ''etc.'')
***** M20a3 - Madagascar
Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
, USA (New Jersey
New Jersey is a U.S. state, state located in both the Mid-Atlantic States, Mid-Atlantic and Northeastern United States, Northeastern regions of the United States. Located at the geographic hub of the urban area, heavily urbanized Northeas ...
)
*** M51 – in Cambodia
Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
and Indonesia
**** M51a
***** M51a1
****** M51a1a
****** M51a1b
***** M51a2
**** M51b
***** M51b1
****** M51b1a
****** M51b1b
** Haplogroup M
– found in South Asia, with highest concentrations in SE India and Bangladesh; oldest haplogroup M lineage on the Indian sub-continent. Also found with low frequency in Southwest China, southwestern China.
*** M2a'b
**** M2a – India (Madhya Pradesh), Munda; most common in Bangladesh
***** M2a1 – Malpaharia
****** M2a1a – Uyghur, Sindhi, Hill Kolam, Thailand
******* M2a1a1 – Katkari
******** M2a1a1a – Nihal
********* M2a1a1a1 – Katkari
******** M2a1a1b – Nihal
********* M2a1a1b1 – Nihal
******* M2a1a2 – Madia
******** M2a1a2a – Madia
********* M2a1a2a1 – Kamar
********** M2a1a2a1a – Kamar
******* M2a1a-G207A
******** M2a1a3 – Mathakur
********* M2a1a3-T16093C
********** M2a1a3a – Kathakur, Mathakur
*********** M2a1a3a1 – Kathakur
********** M2a1a3b – Kathakur
****** M2a1b – Dungri Bhil
****** M2a1c – Andh
***** M2a2
****** M2a2a
***** M2a3
****** M2a3a
**** M2b – Paudi Bhuiya; most common in SE India
***** M2b1 – Korku
****** M2b1a – Korku, Munda
****** M2b1b – Malpaharia
***** M2b2 – Hill Kolam, Jenu Kuruba
***** M2b3 – Betta Kurumba
****** M2b3a – Betta Kurumba
***** M2b4 – Korku
*** M2c – Myanmar, Thailand/Laos
** Haplogroup M3 – Uyghur, Myanmar, New Delhi (Hindu), Paniy
– found mainly in South Asia, with highest concentrations in west and NW India
*** M3a
**** M3a1 – Dongri Bhil, Kathodi, Katkari, Jammu and Kashmir, Pathan, Iran, Thailand/Laos
***** M3a1-T204C
****** M3a1a – Kamar of Chhattisgarh
****** M3a1b
******* M3a1b* – Tajiks in Xinjiang, Sarikolis and Wakhis in Taxkorgan, Pakistan (Balochi, Makrani), India
******* M3a1b1
******** M3a1b1* – Jammu and Kashmir
******** M3a1b1a – Pakistan
Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
( Hazara)
******* M3a1b2 – Pakistan
Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
( Brahui), Iran
Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...
( Persian)
**** M3a2 – Bangladesh, Jammu and Kashmir, Burusho, Qatar, Yemen
***** M3a2a – Jenu Kuruba
*** M3b – Kamar of Chhattisgarh
*** M3c – Madia, Myanmar
**** M3c-T152C!
***** M3c1
****** M3c1a – Jammu and Kashmir, Nepal (Terai Hindu, Tharu), Andhra Pradesh (tribal)
****** M3c1b – Hill Kolam
******* M3c1b1 – Saudi Arabia
******** M3c1b1a – Jenu Kuruba
******** M3c1b1b – Jenu Kuruba
**** M3c2 – Pakistan (Brahui), Jammu and Kashmir, Andh, Thailand
*** M3d – Nepal (Kathmandu), India, Italy (Salerno)
**** M3d1 – New Delhi (Hindu)
***** M3d1a – Nepal (Kathmandu), Cambodia ( Lao), United Kingdom
****** M3d1a1 – Tibet (Sherpa)
** M4'30 or M4'67
*** M4'67-T16311C!
**** Haplogroup M
– found mainly in South Asia but some sequences in Eastern Saudi Arabia
***** Haplogroup M4a – found in Gujarat
Gujarat () is a States of India, state along the Western India, western coast of India. Its coastline of about is the longest in the country, most of which lies on the Kathiawar peninsula. Gujarat is the List of states and union territories ...
, India
***** Haplogroup M4b – found among ancient specimens in the Euphrates valley
**** Haplogroup M65
***** Haplogroup M65a – found in India, Pakistan (Balochi, Sindhi), Sarikoli in Taxkorgan, Xinjiang, China, Pamiri in Gorno-Badakhshan, Tajikistan, Ladakh, Myanmar, China
****** M65a1
****** M65a-C16311T!!
******* M65a2
***** Haplogroup M65b – found in India and in Pakistan (Balochi)
**** M67
*** Haplogroup M18'38
**** Haplogroup M18 – found among Tharus in southern Nepal
Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
and tribal people in Andhra Pradesh
Andhra Pradesh (ISO 15919, ISO: , , AP) is a States and union territories of India, state on the East Coast of India, east coast of southern India. It is the List of states and union territories of India by area, seventh-largest state and th ...
Haplogroup M18a was also found in Mesolithic Sri Lanka
Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
.
***** M18a
***** M18b
***** M18c
**** Haplogroup M38 – found with high frequency among Tharus from Morang District of southeastern Nepal
Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
and as singletons among Tharus from Chitwan District of south-central Nepal and Hindus
Hindus (; ; also known as Sanātanīs) are people who religiously adhere to Hinduism, also known by its endonym Sanātana Dharma. Jeffery D. Long (2007), A Vision for Hinduism, IB Tauris, , pp. 35–37 Historically, the term has also be ...
from New Delhi
New Delhi (; ) is the Capital city, capital of India and a part of the Delhi, National Capital Territory of Delhi (NCT). New Delhi is the seat of all three branches of the Government of India, hosting the Rashtrapati Bhavan, New Parliament ...
***** M38a
***** M38-T195C!
****** M38b
****** M38c
***** M38-T199C
****** M38d
****** M38e
*** Haplogroup M30 – mainly in India; also found in Nepal, Pakistan, Central Asia (Kyrgyz, Wakhi, and Sarikoli in Taxkorgan, Xinjiang, China and Tajiks in Dushanbe, Tajikistan), the Middle East, and North Africa.
**** M30a
***** M30a1
***** M30a2
**** M30b
**** M30c
***** M30c1
****** M30c1a
******* M30c1a1
**** M30d
***** M30d1
***** M30d2
**** M30-C16234T
***** M30e
**** M30f
**** M30g
*** Haplogroup M37
**** M37-T152C!
***** M37-T152C!-C151T
****** Haplogroup M37a – found in Gujarat, India
******* M37a1
****** M37d
**** M37e
***** M37e2
*** M43
**** M43-T16311C!
***** M43a
****** M43a1
**** M43b
*** M45
**** M45a
*** M54
*** M63
*** M64
*** M66
**** M66a
**** M66b
*Haplogroup M
– found in South Asia
**Haplogroup M5a – found in India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
(Jammu and Kashmir, Madhya Pradesh, Kathakur, Gadaba), Thailand ( Mon in Ratchaburi Province and Lopburi Province), Israel, Kyrgyz in Taxkorgan
***M5a1
****M5a1a – India (incl. Jammu and Kashmir)
****M5a1b – India (Jammu and Kashmir, Dongri Bhil, Nihal, Andh), Pakistan (Burusho), Russia, Spain (Romani), USA (Georgia), USA (California)
***M5a2 – India
****M5a2a – Pakistan (Balochi), India ( Nihal), Thailand ( Tai Yuan in Uttaradit Province)
*****M5a2a1 – India (Hindus in New Delhi), Pakistan (Sindhi)
******M5a2a1a – Saudi Arabia, Iran (Persian), Kazakh, Pakistan (Balochi), India (Dongri Bhil
Bhil or Bheel refer to the various Indigenous peoples, indigenous groups inhabiting western India, including parts of Rajasthan and Madhya Pradesh and are also found in distant places such as Bengal and Tripura. Though they now speak the Bhili ...
, Korku, Lachungpa), Myanmar
*****M5a2a2 – India ( Kamar of Chhattisgarh), Yemen
*****M5a2a3 – India (Pauri Bhuiya, Munda)
*****M5a2a4 – Iran (Persians
Persians ( ), or the Persian people (), are an Iranian ethnic group from West Asia that came from an earlier group called the Proto-Iranians, which likely split from the Indo-Iranians in 1800 BCE from either Afghanistan or Central Asia. They ...
), Pakistan (Brahui, Makrani, Balochi)
***M5a3
****M5a3a – India ( Kamar of Chhattisgarh)
****M5a3b – India (Dongri Bhil, Kathodi)
***M5a4 – India (Kathodi, Korku)
***M5a5 – India (Dongri Bhil, Andh), Yemen
**Haplogroup M5b – found in India and Thailand ( Khon Mueang in Chiang Mai Province)
***Haplogroup M5b2b1a – found in Tibet, Ladakh, Nepal
**Haplogroup M5c – found in India, Thailand ( Mon in Lopburi Province and Nakhon Ratchasima Province), Tibet, Nepal
***Haplogroup M5c1 – India (Pauri Bhuiya, Kathodi, ''etc.''), Thailand ( Thai from Phichit Province[Kutanan,W., Kampuansai,J., Brunelli,A., Ghirotto,S., Pittayaporn,P., Ruangchai,S., Schroder,R., Macholdt,E., Srikummool,M., Kangwanpong,D., Hubner,A., Arias,L. and Stoneking,M., "New insights from Thailand into the maternal genetic history of Mainland Southeast Asia." ''Eur. J. Hum. Genet.'' (2018).])
***Haplogroup M5c2 – Nepal (Tharu), Tibet (Sherpa), Thailand ( Mon from Nakhon Ratchasima Province)
*Haplogroup M
– found mainly in South Asia, with highest concentrations in mid-eastern India and Kashmir
**Haplogroup M6b – found in Kerala
Kerala ( , ) is a States and union territories of India, state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile ...
, India
**Haplogroup M61 – found among ancient specimens in the Euphrates valley
*Haplogroup M
– found in East Asia and Southeast Asia, especially in Japan, southern China, Vietnam
Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
, Laos
Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
, and Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
; also found with low frequency in Central Asia and Siberia
**Haplogroup M7a
***Haplogroup M7a* – Japan
***Haplogroup M7a1
****Haplogroup M7a1* – Japan, Jiangsu, Shandong
****Haplogroup M7a1a
*****Haplogroup M7a1a* – Japan, Korea, Beijing, Hebei, Henan, Sichuan, Shanghai, Shandong
*****Haplogroup M7a1a1
******Haplogroup M7a1a1* – Japan, Korea, Jiangsu, Shandong, Liaoning, Henan
******Haplogroup M7a1a1a – Japan
*****Haplogroup M7a1a2
******Haplogroup M7a1a2* – Japan, Jiangsu
******Haplogroup M7a1a2a – Japan
*****Haplogroup M7a1a3 – Japan
*****Haplogroup M7a1a4
******Haplogroup M7a1a4* – Japan, Zhejiang
******Haplogroup M7a1a4a – Japan
*****Haplogroup M7a1a5
******Haplogroup M7a1a5* – Japan
******Haplogroup M7a1a5a – Japan, Korea, Tianjin
*****Haplogroup M7a1a6
******Haplogroup M7a1a6* – Japan, Philippines, Jiangsu, Shanxi, Shandong
******Haplogroup M7a1a6a – Japan
*****Haplogroup M7a1a7
******Haplogroup M7a1a7* – Japan, Korea
******Haplogroup M7a1a7a – Uyghur
*****Haplogroup M7a1a8 – Japan, Jiangsu
*****Haplogroup M7a1a9 – Japan, Korea, Tianjin
*****Haplogroup M7a1a10 – Japan
****Haplogroup M7a1b
*****Haplogroup M7a1b1
******Haplogroup M7a1b1* – Japan, China (Minnan Han)
******Haplogroup M7a1b1a – Japan
*****Haplogroup M7a1b2 – Japan
***Haplogroup M7a2
****Haplogroup M7a2* – Japan
****Haplogroup M7a2a – Japan, Ulchi, Yakut
*****Haplogroup M7a2a1 – Japan
*****Haplogroup M7a2a2
******Haplogroup M7a2a2* – Japan
******Haplogroup M7a2a2a
*******Haplogroup M7a2a2a* – Japan ( Gunma)
*******Haplogroup M7a2a2a1 – Japan ( Aichi)
*****Haplogroup M7a2a3
******Haplogroup M7a2a3a
*******Haplogroup M7a2a3a* – Udihe
*******Haplogroup M7a2a3a1 – Udihe
******Haplogroup M7a2a3b – Evenk ( Nyukzha River basin), Buryat
*****Haplogroup M7a2a4 – Japan
**Haplogroup M7b'c
***Haplogroup M7b
****Haplogroup M7b1a
*****Haplogroup M7b1a1 – Thailand, Laos, Vietnam, Cambodia, Myanmar, Indonesia, China, Karakalpak, Kyrgyz, Mongush, Khamnigan
******Haplogroup M7b1a1a – Thailand, Uyghur, Korea
*******Haplogroup M7b1a1a1 – Japan, Korea, China, Tajikistan, Thailand, Laos
********Haplogroup M7b1a1a1a – Japan
********Haplogroup M7b1a1a1b – Japan, Korea, China, Russia, Kyrgyzstan
********Haplogroup M7b1a1a1c – Japan
********Haplogroup M7b1a1a1d – Japan
*******Haplogroup M7b1a1a2 – Thailand, Vietnam, Malaysia, China
*******Haplogroup M7b1a1a3 – Thailand, Laos, Vietnam, China
******Haplogroup M7b1a1b – Thailand, Laos, Vietnam, China (Hunan, Uyghur, Kyrgyz from Artux), England
******Haplogroup M7b1a1c – Han Chinese, Uyghurs, Kyrgyz
*******Haplogroup M7b1a1c1 – Chinese, Bama Yao Autonomous County
Bama Yao Autonomous County ( Zhuang: ,) is a county in Guangxi, China. It is under the administration of Hechi City. The residents of Bama County have a reputation for longevity, and Bama has been the focus of studies from geriatricians nationwid ...
******Haplogroup M7b1a1d – Thailand, Laos, Tatar (Buinsk)
******Haplogroup M7b1a1e – Thailand
*******Haplogroup M7b1a1e1 – Thailand, Vietnam, China
*******Haplogroup M7b1a1e2 – China
******Haplogroup M7b1a1f – Thailand, Malaysia, Indonesia, Vietnam, China
******Haplogroup M7b1a1g – Thailand
******Haplogroup M7b1a1h – Thailand, Chinese ( Han from Lanzhou, ''etc.''), Vietnam, Korea, Japan
******Haplogroup M7b1a1i – Taiwan ( Amis), Philippines, Malaysia
*****Haplogroup M7b1a2
******Haplogroup M7b1a2a – China (Uyghurs, Kyrgyzes in Taxkorgan, Han, Mongol in Inner Mongolia)
*******Haplogroup M7b1a2a1 – Taiwan Aboriginal peoples, Philippines, Indonesia, Malaysia
********Haplogroup M7b1a2a1a – Atayal, Saisiyat
********Haplogroup M7b1a2a1b – Atayal
*********Haplogroup M7b1a2a1b1 – Atayal, Saisiyat
****Haplogroup M7b1b – Khamnigan, China, Kyrgyz
***Haplogroup M7c
****Haplogroup M7c1 – China, Vietnam, Malaysia, Mongolia, Sarikoli, Kazakhstan
*****Haplogroup M7c1a – China, Korea, Japan, Vietnam, Thailand, Indonesia
******Haplogroup M7c1a1
*******Haplogroup M7c1a1a – China, Mongolia
*******Haplogroup M7c1a1b – Azeri
********Haplogroup M7c1a1b1 – Even ( Sakkyryyr), Yakut ( Vilyuy basin)
******Haplogroup M7c1a2 – China
*******Haplogroup M7c1a2a – She, Uyghur
********Haplogroup M7c1a2a1 – Japan, Korea, Uyghur
******Haplogroup M7c1a3 – China, Japan, Vietnam
*******Haplogroup M7c1a3a – Korea
******Haplogroup M7c1a4
*******Haplogroup M7c1a4a – China
*******Haplogroup M7c1a4b – China
******Haplogroup M7c1a5 – Japan, Korea
*****Haplogroup M7c1b – Chinese
******Haplogroup M7c1b1 – Buryats
******Haplogroup M7c1b2
*******Haplogroup M7c1b2a – Khamnigan, Korea
*******Haplogroup M7c1b2b – China, Thailand, Laos, Malaysia
*****Haplogroup M7c1c – China, Thailand/Laos
******Haplogroup M7c1c1 – China
*******Haplogroup M7c1c1a – China
********Haplogroup M7c1c1a1 – Philippines
******Haplogroup M7c1c2 – Thailand, China (Han)
******Haplogroup M7c1c3 – Taiwan, Thailand, Philippines, Indonesia, Brunei, Malaysia, Kiribati, Nauru, Saudi Arabia, Madagascar
****Haplogroup M7c2 – Taiwan, Hainan, Thailand/Laos
*****Haplogroup M7c2a – Thailand/Laos, China (incl. Hainan)
*****Haplogroup M7c2b – Thailand, Taiwan (Han), Czech
****Haplogroup M7c3 – China (incl. Amis)
*Haplogroup M8 - China, Northern Thailand ( Lisu), India
**Haplogroup M8a
– found in East Asia, Central Asia, and Siberia
***Haplogroup M8a1
****Haplogroup M8a1a – Japan
Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
****Haplogroup M8a1b – southeastern Siberia ( Udegey)
***Haplogroup M8a2'3
****Haplogroup M8a2'3* – Japan
****Haplogroup M8a2 – found in Koryaks, Itelmens, Chukchis, Tuvans, Khakassians, Altayans, Mongolians, China (including Uyghurs
The Uyghurs,. alternatively spelled Uighurs, Uygurs or Uigurs, are a Turkic peoples, Turkic ethnic group originating from and culturally affiliated with the general region of Central Asia and East Asia. The Uyghurs are recognized as the ti ...
), Koreans
Koreans are an East Asian ethnic group native to the Korean Peninsula. The majority of Koreans live in the two Korean sovereign states of North and South Korea, which are collectively referred to as Korea. As of 2021, an estimated 7.3 m ...
, Japan, Thailand/Laos
*****Haplogroup M8a2* – China (Hakka)
*****Haplogroup M8a2-T152C!!!
******Haplogroup M8a2-T152C!!!* – China, Japan (Chiba)
******Haplogroup M8a2a
*******Haplogroup M8a2a1 – found in Thailand, China, Japan
********Haplogroup M8a2a1a
*********Haplogroup M8a2a1a* – Northeast Thailand ( Saek)
*********Haplogroup M8a2a1a1
**********Haplogroup M8a2a1a1* – China ( Han from Wuhan
Wuhan; is the capital of Hubei, China. With a population of over eleven million, it is the most populous city in Hubei and the List of cities in China by population, eighth-most-populous city in China. It is also one of the nine National cent ...
)
**********Haplogroup M8a2a1a1a – Central Thailand (Tai Yuan), Northern Thailand ( Palaung)
********Haplogroup M8a2a1b
*********Haplogroup M8a2a1b* – Chiang Mai Province ( Khon Mueang)
*********Haplogroup M8a2a1b1 – Lamphun Province ( Khon Mueang)
********Haplogroup M8a2a1c – China, Japan ( Aichi)
*******Haplogroup M8a2a2 – China
******Haplogroup M8a2-A12530G/G14364A/T16297C – Uyghur
******Haplogroup M8a2b – China ( Maonan from Pingtang County)
*******Haplogroup M8a2b* - Japan, China (Shandong, Henan)
*******Haplogroup M8a2b1 (C9301T) - China (Anhui Medical University Hospital, ''etc.'')
*******Haplogroup M8a2b2 (G12192A) - Russia ( Ulchi, Nanai from Bogorodskoye, Ulchsky District)
*******Haplogroup M8a2b3 (T16311C!) - China, Thailand (Bangkok)
*******Haplogroup M8a2b4 (T9813d) - China (Zhejiang, ''etc.'')
*****Haplogroup M8a2c – found in Japan and China
*****Haplogroup M8a2d – found in China (Shantou, Qingdao)
*****Haplogroup M8a2e – found in Taiwan ( Ami, ''etc.'') and in a Han Chinese living in the Denver, Colorado metropolitan area
*****Haplogroup M8a2f – China
****Haplogroup M8a3
*****Haplogroup M8a3* – China (Guangdong
) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
, ''etc.''), Kyrgyz ( Artux), Russia (Verkhnyaya Gutara, Nerkha, and Kushun villages of Irkutsk Oblast
Irkutsk Oblast (; ) is a federal subjects of Russia, federal subject of Russia (an oblast), located in southeastern Siberia in the basins of the Angara River, Angara, Lena River, Lena, and Nizhnyaya Tunguska Rivers. The administrative center is ...
)
*****Haplogroup M8a3a
******Haplogroup M8a3a* – China, Russia ( Ket from Turukhansk)
******Haplogroup M8a3a1
*******Haplogroup M8a3a1* – China
*******Haplogroup M8a3a1a – China
******Haplogroup M8a3a2 – China, Indonesia (Jawa Timur)
** Haplogroup CZ - Northern Thailand ( Hmong)
*** Haplogroup Cbr>
– found especially in Siberia
****Haplogroup C
– found in Asia and America ( Indigenous peoples of the Americas, Native Americans and Hispanics in particular)
****Haplogroup C4
****Haplogroup C
– found in China and Ukraine
*** Haplogroup Z (mtDNA), Haplogroup Zbr>
– found in Northeast Europe, Siberia, Central Asia, and East Asia, including among Swedes
Swedes (), or Swedish people, are an ethnic group native to Sweden, who share a common ancestry, Culture of Sweden, culture, History of Sweden, history, and Swedish language, language. They mostly inhabit Sweden and the other Nordic countries, ...
, Sami
Acronyms
* SAMI, ''Synchronized Accessible Media Interchange'', a closed-captioning format developed by Microsoft
* Saudi Arabian Military Industries, a government-owned defence company
* South African Malaria Initiative, a virtual expertise ne ...
, Finns
Finns or Finnish people (, ) are a Baltic Finns, Baltic Finnic ethnic group native to Finland. Finns are traditionally divided into smaller regional groups that span several countries adjacent to Finland, both those who are native to these cou ...
, Russians
Russians ( ) are an East Slavs, East Slavic ethnic group native to Eastern Europe. Their mother tongue is Russian language, Russian, the most spoken Slavic languages, Slavic language. The majority of Russians adhere to Eastern Orthodox Church ...
, Ukrainians, Nogais, Abazins, Cherkessians, North Ossetians, Turks, Udmurts, Komi, Kets, Kalmyks, Hazara, Pashtuns, Tajiks, Turkmens, Uzbeks, Kazakhs, Kyrgyz, Uyghurs, Evens, Evenks, Dolgans, Yakuts, Yukaghirs, Khakas, Altaians, Altai Kizhi, Buryats, Nganasans, Koryaks, Itelmens, Ulchi, Japanese, Koreans
Koreans are an East Asian ethnic group native to the Korean Peninsula. The majority of Koreans live in the two Korean sovereign states of North and South Korea, which are collectively referred to as Korea. As of 2021, an estimated 7.3 m ...
, Chinese, and Tibeto-Burman peoples
*Haplogroup M
– found in East Asia and Central Asia, especially in Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
. In the Nepalese populations, it is prevalent mainly in Sherpa (27.4%), Tharu-CI (19.6%), Tamang (15.5%), Magar (13.5%), and Tharu-CII. Haplogroup M9* has additionally been found in ancient remains from the Red Deer Cave people in present-day Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
.
**Haplogroup M9a'b
***Haplogroup M9a – Han (Guangdong, Guangxi, Yunnan, Sichuan, Hunan, Taiwan, Anhui, Shaanxi, Shandong, Hebei), Korean (South Korea), Tujia (Hunan), Kinh (Hue), Mongol (Hohhot), Japanese, Lhoba 28,800) ybp">MRCA 23,000 (95% CI 18,100 <-> 28,800) ybp****Haplogroup M9a1 – Han (Hunan) 26,700) ybp">MRCA 19,500 (95% CI 13,800 <-> 26,700) ybp*****Haplogroup M9a1a – Han (Hebei, Henan, Shaanxi, Anhui, Zhejiang, Hunan, Yunnan, Guangdong, Hong Kong, Taiwan), Manchu (Jilin), Korean (South Korea), Hui (Qinghai), Kazakh (Ili), Kyrgyz (Kyrgyzstan), Nepal 20,900) ybp">MRCA 16,500 (95% CI 12,800 <-> 20,900) ybp******Haplogroup M9a1a1 – Han (Henan, Shaanxi, Guangdong, Guangxi, Sichuan, Yunnan), Taiwan, Thailand/Laos, Hui ( Yuxi), Tibetan (Nyingchi), Uyghur, Japanese (Hokkaido) 17,600) ybp">MRCA 13,900 (95% CI 10,800 <-> 17,600) ybp*******Haplogroup M9a1a1a – Japanese, Korean (Seoul), Chinese (incl. a Henan Han), Khamnigan (Buryat Republic), Udege, Nivkh, Tibetan (Qinghai)
*******Haplogroup M9a1a1b – Japanese, Korean (South Korea), Mongol (Inner Mongolia), Han (Hunan)
*******Haplogroup M9a1a1c – Han (Gansu, Shaanxi, Henan, Liaoning, Zhejiang, Jiangxi, Hunan, Guangdong, Sichuan, Yunnan), Ainu, Japanese, Korean, Mongol (Hohhot), Uyghur (Ürümqi), Altaian, Tuvinian, Hui (Xinjiang, Kyrgyzstan), Tujia (Hunan), Bai (Yunnan), Yi (Yunnan)
********Haplogroup M9a1a1c1 – Han (Henan)
*********Haplogroup M9a1a1c1a – Han (Henan, Anhui, Shandong, Liaoning, Sichuan, Yunnan, Xinjiang, Lanzhou), Korea, Japanese, Mongol (New Barga Left Banner), Tibetan (Liangshan), Hui (Ili)
*********Haplogroup M9a1a1c1b – Tibetan (Gansu, Qinghai, Sichuan, Yunnan, Chamdo, Lhasa, Nagqu, Ngari, Nyingchi, Shannan, Shigatse), Monpa (Nyingchi), Dirang Monpa (Arunachal Pradesh), Lachungpa (Sikkim), Tu (Huzhu Tu Autonomous County), Dongxiang (Gansu), Buryat (Inner Mongolia, Buryat Republic), Han (Qinghai), Hui (Qinghai), Nepalese
*******Haplogroup M9a1a1d – Salar (Qinghai), Han ( Yanting), Bai (Dali)
******Haplogroup M9a1a2 – Tharu (Chitwan District, Uttar Pradesh), Tibetan (Nagqu, Yunnan, Qinghai, Shigatse), Lhoba (Nyingchi), Dhimal (West Bengal), Chin (Myanmar), Adi (Assam), Tu (Qinghai), Uyghur (Ürümqi), Mongol (Ili), Han (Hunan, Shanxi, Sichuan, Yunnan, Shandong, Ili), Yi (Yunnan), Bai (Dali), Nepalese Behar et al., 2012b">MRCA 6,153.9 ± 5,443.2 ybp; CI=95%[Behar et al., 2012b]*****Haplogroup M9a1b – Tibetan (Nyingchi, Nagqu, Lhasa, Chamdo, Ngari, Shannan, Shigatse, Sichuan, Yunnan, Qinghai, Gansu), Monba (Nyingchi), Lhoba (Shannan), Uzbekistan (Fergana), Dongxiang (Linxia), Naga (Sagaing), Burman (Bago), Chin (Chin State), Han (Hunan, Sichuan, Yunnan, Liaoning), Yi (Shuangbai) ">MRCA 9,416.6 ± 3,984.0 ybp; CI=95%******Haplogroup M9a1b1 – Tibetan, Lhoba, Arunachal Pradesh (Sonowal Kachari, Wanchoo, Gallong), Assam (Adi), Sikkim (Lepcha, Lachung), Qinghai (Salar, Tu), Mongol (Mongolia, Inner Mongolia), Guangxi (Gelao, Palyu), Thailand, Bengal, Pakistan (Karachi), Meghalaya (Khasi, Garo), Bodo (West Bengal), Rabha (West Bengal), Rajbanshi (West Bengal), Indonesia, Han (Shaanxi, Henan, Gansu, Sichuan, Yunnan), Hui (Gansu, Qinghai), Burman (Ayeyarwady, Magway, Sagaing), Rakhine (Rakhine, Magway), Chin (Chin State), Naga (Sagaing), Mech (Jhapa district, Nepal), Nepalese, Mosuo (Yunnan), Yi (Yunnan), She (Guizhou), Hani (Yunnan), Pumi (Yunnan), Bai (Dali), Va (Yunnan) ">MRCA 6,557.4 ± 2,102.4 ybp; CI=95%******Haplogroup M9a1b2 – Tibetan (Diqing), Han (Dujiangyan), Kazakh (Altai Republic), Kalmyk ">MRCA 3,225.9 ± 3,494.4 ybp; CI=95%****Haplogroup M9a4
*****Haplogroup M9a4a – Kinh (Hanoi), Han (Shaanxi, Shandong, Zhejiang, Taiwan, Sichuan, Guangdong), Li (Hainan), Mulam (Guangxi), Jino (Xishuangbanna), Dai (Xishuangbanna), Chiang Mai
*****Haplogroup M9a4b – Kinh (Hanoi), South Korea
****Haplogroup M9a5 – Han (Hunan, Hong Kong), Thailand, Pubiao ( Malipo), Li (Hainan), Mulam ( Luocheng), Zhuang (Bama Yao Autonomous County
Bama Yao Autonomous County ( Zhuang: ,) is a county in Guangxi, China. It is under the administration of Hechi City. The residents of Bama County have a reputation for longevity, and Bama has been the focus of studies from geriatricians nationwid ...
), Kinh (Hanoi)
***Haplogroup M9b – Han (Luocheng, Dujiangyan, Shaanxi), Cham (Binh Thuan), Mulam (Luocheng), Bouyei (Guizhou), Yi (Hezhang), Bunu (Dahua), Hui (Ili), Thailand ( Phuan from Sukhothai Province)
** Haplogroup E – a subclade of M9 – found especially in Taiwan (Aboriginal peoples), Maritime Southeast Asia, and the Mariana Islands ">MRCA 23,695.4 ± 6,902.4 ybp; CI=95%*Haplogroup M1
– small clade found in East Asia, Southeast Asia, Bangladesh, Central Asia, Saudi Arabia, southern Siberia, Russia, Belarus, and Poland 31,700) ybp">MRCA 23,600 (95% CI 17,100 <-> 31,700) ybp**M10-514C!/A15218G/C16362T!
***M10-514C!/A15218G/C16362T!* – Poland
***M10-T3167C/C4140T/T8793C/C12549T/A13152G/T14502C/C15040T/T15071C
****M10a 22,800) ybp">MRCA 16,700 (95% CI 11,800 <-> 22,800) ybp*****M10a* – Myanmar
*****M10a1 – China, Thailand, Myanmar, Japan, Shor, Daur 18,400) ybp">MRCA 14,400 (95% CI 11,100 <-> 18,400) ybp******M10a1* – Myanmar, Central Thailand ( Mon), Japan (Aichi)
******M10a1-G16129A!!!
*******M10a1-A13105G!/T16362C – Shor, Daur
*******M10a1a
********M10a1a* – China, Saudi Arabia
********M10a1a1
*********M10a1a1a – Mongolia, Korea, Japan, China (Han from Kunming, ''etc.''), Iron Age Black Sea Scythian
*********M10a1a1b – Altai, Korea, Japan, China
**********M10a1a1b* – Chinese Uyghur
**********M10a1a1b1
***********M10a1a1b1* – Japan, China
***********M10a1a1b1a – China
***********M10a1a1b1b – China
***********M10a1a1b1c – Uyghur, Altai-Kizhi
**********M10a1a1b2
***********M10a1a1b2* – Taiwan (Hakka)
***********M10a1a1b2a – Japan
********M10a1a2 – Northern Thailand (Khon Mueang from Lamphun Province)
********M10a1a3 – Taiwan
*******M10a1b
********M10a1b* – South Korea, China (Han from Tai'an, ''etc.''), India ( Gallong, ''etc.'')
********M10a1b1 – China ( Makatao)
********M10a1b2 – China ( Tingri County)
*******M10a1c – China
*****M10a2 – Japan (Aichi), Kalmyk, Russian (northwestern Russia)
**M10b – China ( Shui), Vietnam ( Cờ Lao)
*Haplogroup M1
– small clade found especially among the Han Chinese, Chinese and also in some Japanese, Koreans
Koreans are an East Asian ethnic group native to the Korean Peninsula. The majority of Koreans live in the two Korean sovereign states of North and South Korea, which are collectively referred to as Korea. As of 2021, an estimated 7.3 m ...
, Oroqen, Yi, Tibetans, Tajiks in Dushanbe, Tajikistan, and Bangladeshis ">MRCA 20,987.7 ± 5,740.8 ybp; CI=95%**Haplogroup M11* – Myanmar
**Haplogroup M11a'b'd
***Haplogroup M11a'b ">MRCA 16,209.0 ± 4,396.8 ybp; CI=95%****Haplogroup M11a – Korea, Turkey ">MRCA 11,972.3 ± 3,523.2 ybp; CI=95%*****Haplogroup M11a-C198T
******Haplogroup M11a-C198T* – Wancho, Miao (from Fenghuang, Hunan)
******Haplogroup M11a1 – Gallong, Tibet ">MRCA 8,668.6 ± 4,041.6 ybp; CI=95%*****Haplogroup M11a2 – Tibet, Han (Zhanjiang) ">MRCA 8,776.3 ± 3,715.2 ybp; CI=95%*****Haplogroup M11a3 – Uyghur, Buryat, Oroqen
****Haplogroup M11b ">MRCA 12,962.0 ± 4,819.2 ybp; CI=95%*****Haplogroup M11b1 – Taiwan (Minnan)
******Haplogroup M11b1a – Japan
*******Haplogroup M11b1a1 – Japan, Han (Tai'an)
*****Haplogroup M11b2 – Japanese (Hokkaido), China, Altai-Kizhi, Tajik (Dushanbe)
***Haplogroup M11d – China, Teleut, Kyrgyz, Iran
**Haplogroup M11c – Japan, Korea
*Haplogroup M12'G
**Haplogroup M1
– small clade found especially among the Li people, aborigines of Hainan Island as well as in other populations of China, Japan, Korea, Pashtuns
Pashtuns (, , ; ;), also known as Pakhtuns, or Pathans, are an Iranian peoples, Iranic ethnic group primarily residing in southern and eastern Afghanistan and northwestern Pakistan. They were historically also referred to as Afghan (ethnon ...
, Tibet, Myanmar, Thailand, Cambodia, and Vietnam ">MRCA 31,287.5 ± 5,731.2 ybp; CI=95%***Haplogroup M12a ">MRCA 26,020.6 ± 5,808.0 ybp; CI=95%****Haplogroup M12a1 – Thailand/Laos
*****Haplogroup M12a1a – Thailand ( Htin in Phayao Province, Black Tai in Kanchanaburi Province, Mon in Kanchanaburi Province, Khon Mueang in Chiang Mai Province), Laos ( Lao in Luang Prabang), Hainan
******Haplogroup M12a1a1 – China (esp. Hainan)
******Haplogroup M12a1a2 – Hainan, Tu
*****Haplogroup M12a1b – Tibet, Thailand ( Blang in Chiang Rai Province, Khon Mueang in Chiang Rai Province, Palaung in Chiang Mai Province, Mon in Kanchanaburi Province), Hainan, Vietnam
****Haplogroup M12a2 – Thailand, Hainan, Myanmar
***Haplogroup M12b – Thailand ( Khmu in Nan Province)
****Haplogroup M12b1 – Vietnam, Myanmar
*****Haplogroup M12b1a – Laos ( Lao in Vientiane)
******Haplogroup M12b1a1
******Haplogroup M12b1a2 – Thailand (Soa in Sakon Nakhon Province)
*******Haplogroup M12b1a2a – Cambodia, Malaysia
*******Haplogroup M12b1a2b – Cambodia
*****Haplogroup M12b1b – Thailand ( Suay in Surin Province, Khmer in Surin Province, Lao Isan in Roi Et Province, Black Tai in Loei Province), Cambodia
****Haplogroup M12b2 – Thailand, Hainan
*****Haplogroup M12b2a – Cambodia
** Haplogroup Gbr>
– found especially in Japan, Mongolia, and Tibet and in indigenous peoples of Kamchatka Peninsula, Kamchatka ( Koryaks, Alyutors, Itelmens), with some isolated instances in diverse places of Asia ">MRCA 31,614.8 ± 5,193.6 ybp; CI=95%***Haplogroup G1 – Japan ">MRCA 21,492.9 ± 5,414.4 ybp; CI=95%****Haplogroup G1a – China (Uyghurs), Thailand ( Black Lahu in Mae Hong Son Province) ">MRCA 18,139.1 ± 5,462.4 ybp; CI=95%(TMRCA 18,800 26,900">5% CI 12,600 <-> 26,900ybp)
*****Haplogroup G1a1 – Korea, Vietnam ( Dao), China (Sarikolis, Uyghurs, ''etc.''), Tajikistan (Pamiris), Russia ( Todzhin) 14,600) ybp">MRCA 12,200 (95% CI 10,100 <-> 14,600) ybpref name = "YFull" />
******Haplogroup G1a1a – Japan, Korea, Taiwan 7,000) ybp">MRCA 5,200 (95% CI 3,800 <-> 7,000) ybpref name = "YFull" />
*******Haplogroup G1a1a1 – Japan, Korea, China ( Daur, Korean in Arun Banner) (TMRCA 3,100 6,300">5% CI 1,250 <-> 6,300ybp)
*******Haplogroup G1a1a2 – Japan
*******Haplogroup G1a1a3 – Japan
*******Haplogroup G1a1a4 – Japan, Korea
******Haplogroup G1a1b – Taiwan ( Makatao), Russia ( Altai-Kizhi)
*****Haplogroup G1a2'3
******Haplogroup G1a2 – Manchu, Han (Beijing), Tibet, Miao
******Haplogroup G1a3 – Japan, Korea ">MRCA 6,451.1 ± 4,521.6 ybp; CI=95%****Haplogroup G1b ">MRCA 7,246.3 ± 5,088.0 ybp; CI=95%(TMRCA 10,300 14,700">5% CI 7,000 <-> 14,700ybp)
*****Haplogroup G1b1 – (TMRCA 8,400 11,100">5% CI 6,200 <-> 11,100ybp) Koryaks ( Magadan Oblast, Severo-Evensk District of Magadan Oblast), Nivkhs, Evens (Kamchatka), Yakut (HGDP)
*****Haplogroup G1b2'3'4 (G1b-G16129A!) – (TMRCA 7,700 11,400">5% CI 5,000 <-> 11,400ybp) Evenks ( Iengra), Evens ( Magadan region), Orok, Nivkhs, Koryaks ( Magadan Oblast, Severo-Evensk District of Magadan Oblast, Kamchatka)
******Haplogroup G1b2 – (TMRCA 2,600 9,300">5% CI 375 <-> 9,300ybp) Koryaks ( Magadan region, Severo-Evensk District of Magadan Region), Chukchi
******Haplogroup G1b3 – (TMRCA 4,700 13,500">5% CI 1,000 <-> 13,500ybp) Chukchi (Anadyr), Evens (Kamchatka)
******Haplogroup G1b4 – (TMRCA 1,450 5,100">5% CI 225 <-> 5,100ybp) Yukaghirs, Even ( Tompo)
****Haplogroup G1c – Korea (Seoul), China ( Han from Lanzhou, ''etc.''), Thailand ( Black and Red Lahu in Mae Hong Son Province), Malaysia ( Seletar) ">MRCA 12,910.6 ± 6,009.6 ybp; CI=95%(TMRCA 17,200 24,600">5% CI 11,600 <-> 24,600ybp)
*****Haplogroup G1c1 – Han (Sichuan
Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
, Tai'an)
*****Haplogroup G1c2 – China (including at least one Han from Beijing)
***Haplogroup G2 – Cambodia ">MRCA 26,787.5 ± 4,617.6 ybp; CI=95%****Haplogroup G2a'c – China, USA
*****Haplogroup G2a ">MRCA 17,145.5 ± 5,270.4 ybp; CI=95%******Haplogroup G2a1 – Korea, China (Hebei
Hebei is a Provinces of China, province in North China. It is China's List of Chinese administrative divisions by population, sixth-most populous province, with a population of over 75 million people. Shijiazhuang is the capital city. It bor ...
, Eastern China), Tibetans ( Chamdo, Lhasa
Lhasa, officially the Chengguan District of Lhasa City, is the inner urban district of Lhasa (city), Lhasa City, Tibet Autonomous Region, Southwestern China.
Lhasa is the second most populous urban area on the Tibetan Plateau after Xining ...
, Tingri), Uyghur ( Artux), Daur, India (Ladakh, Punjabi Hindu), Myanmar ( Bamar from Kayin State
Kayin State (, ; ; , ), formerly known as Karen State, is a Administrative divisions of Myanmar, state of Myanmar. The capital city is Hpa-An, also spelled Pa-An.
The terrain of the state is mountainous; with the Dawna Range running along the ...
), Thailand, Singapore, Kyrgyz, Kazakhstan, Afghanistan (Balkh), Israel ( haMerkaz), Saudi Arabia, Russia (Chelyabinsk Oblast
Chelyabinsk Oblast; , is a federal subjects of Russia, federal subject (an oblast) of Russia in the Ural Mountains region, on the border of Europe and Asia. Its administrative center is the types of inhabited localities in Russia, city of Chel ...
, Taimyr Evenk, Tuvinian), Belarus ( Lipka Tatars), Norway, USA ("caucasian") ">MRCA 14,045.7 ± 4,742.4 ybp; CI=95%*******Haplogroup G2a1a (C12525T) - Georgia ( Abkhaz), Karachay ( Karachay-Cherkess Republic), Yakuts (Central), Buryat (Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
)
*******Haplogroup G2a1b (A12753G)
********Haplogroup G2a1b1 (T146C! * G207A * T8063C * G9266A * A15758G) - Japan
********Haplogroup G2a1b2 (A8832C) - Thailand ( Mon from Central Thailand)
********Haplogroup G2a1b3 (T16126C) - Thailand ( Tai Khuen from North Thailand)
*********Haplogroup G2a1b3a (A7606G) - Thailand ( Thai from Central Thailand)
*******Haplogroup G2a1-T16189C! - China (Han from Yili, Tibetan from Shigatse), Korea, Japan, Vietnam ( Nung), Georgian
********Haplogroup G2a1-T16189C!-A16194G - Japan ( Aichi)
*********Haplogroup G2a1c – Japan, Korea
**********Haplogroup G2a1c1 – Japan
**********Haplogroup G2a1c2 – Japan
********Haplogroup G2a1-T16189C!-C3654T - Eastern China
********Haplogroup G2a1-T16189C!-G16526A - Uyghurs
********Haplogroup G2a1-T16189C!-T15784C - Korea
*********Haplogroup G2a1-T16189C!-T15784C-T7609C - China, India (Lachungpa)
********Haplogroup G2a1d – Kyrgyz ( Artux), Uzbek (Uzbekistan), Poland ">MRCA 7,303.3 ± 3,657.6 ybp; CI=95%*********Haplogroup G2a1d1 – Japan, Hong Kong
**********Haplogroup G2a1d1a – Japan
*********Haplogroup G2a1d2 – Thailand ( Thai from Ratchaburi Province), China (Shandong), India, England
**********Haplogroup G2a1d2a – Thailand ( Tai Yuan from Ratchaburi Province and Chiang Mai Province, Palaung from Chiang Mai Province, Lisu from Mae Hong Son Province), China ( Yao, Eastern China)
**********Haplogroup G2a1d2c - China (Eastern China), Thailand ( Thai from Central Thailand)
*******Haplogroup G2a1e – Japan, Korea
*******Haplogroup G2a1f - China (Taihang area in Henan province)
*******Haplogroup G2a1g – China, Mongols, Karakalpak
*******Haplogroup G2a1-G13194A - Tibetan ( Tingri)
********Haplogroup G2a1h - India (Ladakh, Wancho), China (Fujian
Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
)
*******Haplogroup G2a1i - Lhoba
********Haplogroup G2a1i1 (C3417T) - Yakut ( Vilyuy), Buryat (Eastern Buryat from Buryat Republic)
*******Haplogroup G2a1j (C5840T) - Thailand ( Tai Yuan from North Thailand), China
*******Haplogroup G2a1k (A8521G) - Poland
*******Haplogroup G2a1l (C10043T) - Russia ( Pskov Oblast), Norway (Buskerud
Buskerud () is a Counties of Norway, county and a current electoral district in Norway, bordering Akershus, Oslo, Innlandet, Vestland, Telemark and Vestfold. The region extends from the Oslofjord and Drammensfjorden in the southeast to Hardanger ...
)
*******Haplogroup G2a1m (C8766T) - Buryat (Inner Mongolia)
********Haplogroup G2a1m1 (C7229T) - Daur, Barga (Inner Mongolia), Buryat (Buryat Republic), Khamnigan ( Zabaikal Region)
*********Haplogroup G2a1m1a (A4395G) - Kyrgyzstan ( Kyrgyz)
*******Haplogroup G2a1n (T14180C) - Kyrgyzstan ( Kyrgyz)
*******Haplogroup G2a1o (T173C) - China ( Henan Province)
*******Haplogroup G2a1p (A374G) - Ladakh
*******Haplogroup G2a1q (T16356C) - Deng, China (Fujian
Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
)
******Haplogroup G2a-T152C! - China (Eastern China), Thai ( Chanthaburi)
*******Haplogroup G2a2 – China, Uyghur, Kyrgyz (Artux, Tashkurgan), Kazakhstan, Chelkan (Turochak, Biyka, Kurmach-Baigol), Nogai (Karachay-Cherkess Republic), Yakut ( Vilyuy, Northeast, Central), Buryat ( Kizhinginsky District, Olkhonsky District, Tunkinsky District, Bokhansky District
Bokhansky District (; , ''Bookhonoi aimag'') is an administrative district of Ust-Orda Buryat Okrug of Irkutsk Oblast, Russia, one of the thirty-three in the oblast.Charter of Irkutsk Oblast Municipally, it is incorporated as Bokhansky Municipal ...
), Hazara (Pakistan), Volga Tatar (Republic of Tatarstan), Hungary, Finland
********Haplogroup G2a2a – China, Uyghur, Buryat (Bulagad from Bokhansky District
Bokhansky District (; , ''Bookhonoi aimag'') is an administrative district of Ust-Orda Buryat Okrug of Irkutsk Oblast, Russia, one of the thirty-three in the oblast.Charter of Irkutsk Oblast Municipally, it is incorporated as Bokhansky Municipal ...
), Soyot, Turkey, Bashkortostan, Poland
********Haplogroup G2a2b (G12007A) – Buryat ( Tunkinsky District, Ekhirit-Bulagatsky District)
********Haplogroup G2a2c (T16136C) – Uyghur
********Haplogroup G2a2d (C5456T) - Uyghur
*********Haplogroup G2a2d1 (T8978C) - Uyghur
********Haplogroup G2a2e (T4772C) - Buryat (Ekhirid from Kurumkansky District, Bulagad/Ekhirid from Ekhirit-Bulagatsky District)
*******Haplogroup G2a-T152C!-C16262T - ancient DNA from Irkutsk Oblast
Irkutsk Oblast (; ) is a federal subjects of Russia, federal subject of Russia (an oblast), located in southeastern Siberia in the basins of the Angara River, Angara, Lena River, Lena, and Nizhnyaya Tunguska Rivers. The administrative center is ...
(Irk067 from the Novyj Kachug site at the left bank of the upper Lena River, cal BC 3755 to cal BC 3640)
********Haplogroup G2a3 – Russia (Tatarstan), Azerbaijanian (Iran), Uyghur (Artux)
*********Haplogroup G2a3a – Russia (Tyumen Oblast
Tyumen Oblast () is a federal subjects of Russia, federal subject (an oblast) of Russia. It is located in Western Siberia, and is administratively part of the Ural Federal District. The oblast has administrative jurisdiction over two autonomous ...
, Tatar from Buinsk), Sweden, Germany
*******Haplogroup G2a4 – China (Taihang area in Henan province), Taiwan ( Taitung), Ukraine
********Haplogroup G2a4b (T6707C) - China (Tianjin
Tianjin is a direct-administered municipality in North China, northern China on the shore of the Bohai Sea. It is one of the National Central City, nine national central cities, with a total population of 13,866,009 inhabitants at the time of the ...
, Eastern China)
*******Haplogroup G2a5 – Japan, Korea, Kyrgyz, Kazakh, Karakalpak, Telengit, Tubalar, Yakut, Balkar ( Kabardino-Balkaria), Buryat ( Selenginsky District)
********Haplogroup G2a5a – Buryat ( Barguzinsky District, Kizhinginsky District)
*******Haplogroup G2a-T152C!-T1189C - Uyghur
********Haplogroup G2a-T152C!-T1189C-A10804G - Buryat (Bulagad/Ekhirid from Ekhirit-Bulagatsky District, Bulagad/Ekhirid from Selenginsky District)
********Haplogroup G2a-T152C!-T1189C-C6101T - Daurs
********Haplogroup G2a-T152C!-T1189C-T16271C - China (Liaoning
)
, image_skyline =
, image_alt =
, image_caption = Clockwise: Mukden Palace in Shenyang, Xinghai Square in Dalian, Dalian coast, Yalu River at Dandong
, image_map = Liaoning in China (+all claims hatched).svg
, ...
), Nivkh ( Nogliki)
*******Haplogroup G2a-T152C!-C3351T - China ( Jiangxi Province, Eastern China), Buryat (Ekhirid/Bulagad from Olkhonsky District)
*******Haplogroup G2a-T152C!-C10834T - China (Jilin
)
, image_skyline = Changbaishan Tianchi from western rim.jpg
, image_alt =
, image_caption = View of Heaven Lake
, image_map = Jilin in China (+all claims hatched).svg
, mapsize = 275px
, map_al ...
), Thailand ( Iu Mien from Nan Province)
*****Haplogroup G2-A3397G
******Haplogroup G2-A3397G-T15262C - China, Mongol ( Hulunbuir)
******Haplogroup G2-A3397G-C6623T - Uyghurs
*****Haplogroup G2c – China (Taihang area in Henan province), Uyghurs
****Haplogroup G2b ">MRCA 22,776.4 ± 5,059.2 ybp; CI=95%*****Haplogroup G2b1 – Tibetan ( Shannan), Cambodia ">MRCA 16,830.4 ± 5,616.0 ybp; CI=95%******Haplogroup G2b1a – China (Chongqing
ChongqingPostal Romanization, Previously romanized as Chungking ();. is a direct-administered municipality in Southwestern China. Chongqing is one of the four direct-administered municipalities under the State Council of the People's Republi ...
, Shaanxi
Shaanxi is a Provinces of China, province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to t ...
), Thailand ( Nyaw from Nakhon Phanom Province), Kyrgyzstan, Spain ">MRCA 13,366.8 ± 5,443.2 ybp; CI=95%*******Haplogroup G2b1a1 – Thailand ( Tai Yuan from Uttaradit Province, Khon Mueang from Chiang Mai Province, Karen from Mae Hong Son Province), Myanmar ( Karen from Kayin State
Kayin State (, ; ; , ), formerly known as Karen State, is a Administrative divisions of Myanmar, state of Myanmar. The capital city is Hpa-An, also spelled Pa-An.
The terrain of the state is mountainous; with the Dawna Range running along the ...
and Bago Division
Bago Region (, ; formerly Pegu Division and Bago Division) is an administrative region of Myanmar, located in the southern central part of the country. It is bordered by Magway Region and Mandalay Region to the north; Kayin State, Mon State a ...
)
********Haplogroup G2b1a1a (C16239T) - Thailand ( Skaw Karen from southern Mae Hong Son Province, Skaw Karen from northern Mae Hong Son Province), Myanmar ( Karen from Kayin State
Kayin State (, ; ; , ), formerly known as Karen State, is a Administrative divisions of Myanmar, state of Myanmar. The capital city is Hpa-An, also spelled Pa-An.
The terrain of the state is mountainous; with the Dawna Range running along the ...
)
*******Haplogroup G2b1a2 – Ewenki from Inner Mongolia
Inner Mongolia, officially the Inner Mongolia Autonomous Region, is an Autonomous regions of China, autonomous region of China. Its border includes two-thirds of the length of China's China–Mongolia border, border with the country of Mongolia. ...
********Haplogroup G2b1a2a (G2120A) - China
*******Haplogroup G2b1a3 (A4562T) - Taiwan ( Hakka)
*******Haplogroup G2b1a4 (C4599T) - Thailand ( Tai Khuen from Northern Thailand, Tai Lue from Chiang Rai Province)
*******Haplogroup G2b1a5 (T11353C) - China (Hebei Province), Dungan
******Haplogroup G2b1b – Uyghurs, Tibetan ( Shannan), India (Lachungpa) ">MRCA 4,662.1 ± 4,492.8 ybp; CI=95%*****Haplogroup G2b2 – Late Medieval eastern Mongolia (SHU001), Sweden, Russia, Scotland, Iraq, Japan ">MRCA 20,476.0 ± 5,481.6 ybp; CI=95%******Haplogroup G2b2a – India ( Gallong, Kathodi) ">MRCA 15,328.9 ± 6,057.6 ybp; CI=95%******Haplogroup G2b2b – China, Japan (Chiba)
******Haplogroup G2b2c – China (Taihang area in Henan province), Tatar (Buinsk)
******Haplogroup G2b2d (T5641C) – Japan (Tokyo), Kyrgyz ( Tashkurgan)
*******Haplogroup G2b2d1 (A6647T) - Uyghur, Kumandin ( Soltonsky District), Todzhi (Adir-Kezhig), Buryat (Bulagad/Ekhirid from Ekhirit-Bulagatsky District)
****Haplogroup G2d (A11443G) - Thailand ( Tai Yuan in Central Thailand, Iu Mien in Phayao Province)
*****Haplogroup G2d1 (A328G) - Thailand ( Tai Yuan in Central Thailand, Lisu in Mae Hong Son Province)
***Haplogroup G3 ">MRCA 23,919.9 ± 6,931.2 ybp; CI=95%****Haplogroup G3a
*****Haplogroup G3a1'2 – Pakistan ( Azad Kashmir)
******Haplogroup G3a1 13,000) ybp">MRCA 7,300 (95% CI 3,700 <-> 13,000) ybpref name = "YFull" />
*******Haplogroup G3a1a – Tibetan (Shigatse
Shigatse, officially known as Xigazê () or Rikaze ( zh, s=日喀则, p=Rìkāzé), is a prefecture-level city of the Tibet Autonomous Region of the People's Republic of China. Its area of jurisdiction, with an area of , corresponds to the histo ...
), Sherpa, Naxi, Hani
*******Haplogroup G3a1b – Tibet ( Nagqu), Sarikoli (Tashkurgan)
*******Haplogroup G3a1c (C2389A) - skeletal remains of Bo people from Chang'an Township, Weixin County, Zhaotong, Yunnan, China and similar people from Yangxu Town, Youjiang District, Baise, Guangxi, China and Tham Lod rockshelter, Pang Mapha District, Mae Hong Son Province, Thailand
******Haplogroup G3a1'2-A12661G - Uyghur
******Haplogroup G3a2 30,200) ybp">MRCA 12,600 (95% CI 3,900 <-> 30,200) ybpref name = "YFull" />
*******Haplogroup G3a2* – Taiwan (Minnan)
*******Haplogroup G3a2-T152C!!!
********Haplogroup G3a2-T152C!!!* – Udegey ( Agzu), Korea, Japan
********Haplogroup G3a2a – Japan, Korean, Uzbek
********Haplogroup G3a2b (C13209T) - Uyghur
*********Haplogroup G3a2b1 (G13718C) - China (Eastern China, Shandong)
*****Haplogroup G3a3 24,200) ybp">MRCA 12,900 (95% CI 6,000 <-> 24,200) ybpref name = "YFull" /> – China, Buryat (Khori from Khorinsky District), Ukraine, Poland, Slovakia, Albania, United Kingdom
******Haplogroup G3a3a (G16156A) - China, Uyghur ( Artux)
*******Haplogroup G3a3a1 (A390G) - Inner Mongolia
********Haplogroup G3a3a1a (C2263A) - Uyghur
*********Haplogroup G3a3a1a1 (A3523G) - Bashkortostan
****Haplogroup G3b 24,100) ybp">MRCA 17,800 (95% CI 12,800 <-> 24,100) ybpref name = "YFull" /> - India (Lachungpa), Tibetan (Lhasa
Lhasa, officially the Chengguan District of Lhasa City, is the inner urban district of Lhasa (city), Lhasa City, Tibet Autonomous Region, Southwestern China.
Lhasa is the second most populous urban area on the Tibetan Plateau after Xining ...
), Lhoba
*****Haplogroup G3b1 – Tibet ( Tibetan from Nyingchi, Tibetan from Tingri, Monpa, Deng), India[MtDNA Haplotree at Family Tree DNA](_blank)
/ref> ( Dirang Monpa)
*****Haplogroup G3b2 - China, Tibet ( Monpa, Qinghai), Thailand ( Lawa from the southwest of Mae Hong Son Province, Tai Yuan from Central Thailand)
***Haplogroup G4 – Japan 15,500) ybp">MRCA 7,500 (95% CI 3,100 <-> 15,500) ybpref name = "YFull" />
***Haplogroup G5 – Vietnam ( Dao), Thailand ( Lao Isan from Nakhon Ratchasima Province) 11,200) ybp">MRCA 7,900 (95% CI 5,400 <-> 11,200) ybpref name = "YFull" />
*Haplogroup M13'46'61
**Haplogroup M13 – small clade found among Tibetans in Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
, Oirat Mongols
Mongols are an East Asian ethnic group native to Mongolia, China ( Inner Mongolia and other 11 autonomous territories), as well as the republics of Buryatia and Kalmykia in Russia. The Mongols are the principal member of the large family o ...
in Xinjiang
Xinjiang,; , SASM/GNC romanization, SASM/GNC: Chinese postal romanization, previously romanized as Sinkiang, officially the Xinjiang Uygur Autonomous Region (XUAR), is an Autonomous regions of China, autonomous region of the China, People' ...
, Barghuts in Hulunbuir, Koreans
Koreans are an East Asian ethnic group native to the Korean Peninsula. The majority of Koreans live in the two Korean sovereign states of North and South Korea, which are collectively referred to as Korea. As of 2021, an estimated 7.3 m ...
, and Yakuts and Dolgans in central Siberia ">MRCA 43,169.4 ± 4,944.0 ybp; CI=95%***M13a ">MRCA 19,266.4 ± 6,720.0 ybp; CI=95%****M13a1 – China yghurs, Mongol, Tibetan (Chamdo, Nagqu) India ">MRCA 9,539.2 ± 6,249.6 ybp; CI=95%*****M13a1a – Japan
*****M13a1b – China (Tibet, Uyghurs)
******M13a1b1 – Buryat, Barghut, Yakut (Central), Evenk ( Taimyr)
****M13a2 – Tibet, Thailand ">MRCA 8,726.0 ± 4,828.8 ybp; CI=95%***M13b ">MRCA 31,191.8 ± 6,873.6 ybp; CI=95%****M13b1 – Nepal (Tharu), Northeast India, Thailand, Jahai ">MRCA 22,537.6 ± 7,123.2 ybp; CI=95%****M13b2 – Sherdukpen, Dirang Monpa, Tibet ">MRCA 5,441.4 ± 3,888.0 ybp; CI=95%***M13c – Myanmar, Thailand ( Mon in Kanchanaburi Province), China ( Lahu)
**Haplogroup M46 – Myanmar, Moken, Urak Lawoi, Malagasy
**Haplogroup M61 – Thailand ( Phuan in Suphan Buri Province, Phuan in Phichit Province, Phuan in Sukhothai Province, Saek in Nakhon Phanom Province), Myanmar, Vietnam, Borneo
***Haplogroup M61a – China (Yi, Tibet, Lachungpa)
*Haplogroup M14 – Australia ( Kalumburu), Saudi Arabia, India, Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
?
*Haplogroup M15 – Australia ( Kalumburu), Tibet
Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
?
*Haplogroup M17 – found in Luzon
Luzon ( , ) is the largest and most populous List of islands in the Philippines, island in the Philippines. Located in the northern portion of the List of islands of the Philippines, Philippine archipelago, it is the economic and political ce ...
, Chams, Maniq, Mon, Blang, Lawa, Thai, and Laotians
**Haplogroup M17a – Thailand, Vietnam ( Cham)
**Haplogroup M17c – Vietnam ( Cham)
***Haplogroup M17c1 – Philippines (Abaknon)
****Haplogroup M17c1a – Indonesia
*****Haplogroup M17c1a1 – Philippines (Abaknon)
******Haplogroup M17c1a1a – Cambodia (incl. Stieng)
*Haplogroup M19'53
**Haplogroup M19 – found in the Batak people of Palawan
Palawan (, ), officially the Province of Palawan (; ), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in terms of total area of . The capital and largest c ...
**Haplogroup M53 – India
***Haplogroup M53b – Kamar of Chhattisgarh
*Haplogroup M2
– small clade found in SE Asia ( Semang, Semelai people, Semelai, Temuan, Jehai, Thailand, Maniq, Mon, Karen, Indonesia), China, and Bangladesh
**Haplogroup M21a – Batek
**Haplogroup M21b
***Haplogroup M21b1 – Cambodia ( Lao, Tompoun)
****Haplogroup M21b1a – Semelai people, Semelai, Philippines (Cuyunon of Palawan Island), Indonesia
***Haplogroup M21b2 – Moken, Cham, Malaysia, India
*Haplogroup M22 – Drung (Yunnan)
**Haplogroup M22a – Vietnam (Kinh, Cham), Temuan
**Haplogroup M22b – Vietnam (Kinh), China (Han in Meizhou), Cambodia
*Haplogroup M23'75 – China
**M23 – found in Madagascar
Madagascar, officially the Republic of Madagascar, is an island country that includes the island of Madagascar and numerous smaller peripheral islands. Lying off the southeastern coast of Africa, it is the world's List of islands by area, f ...
, South Africa, USA, Canada
**M75 – found in China, USA
*Haplogroup M24'41
**Haplogroup M24
***Haplogroup M24a – found in Thailand, Cambodia (Khmer), Palawan
Palawan (, ), officially the Province of Palawan (; ), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in terms of total area of . The capital and largest c ...
(Tagbanua), and China
***Haplogroup M24b – found in Thailand, Cambodia (Jarai), and China
**Haplogroup M41 – found in South Asia
***Haplogroup M41b – found in Andhra Pradesh
Andhra Pradesh (ISO 15919, ISO: , , AP) is a States and union territories of India, state on the East Coast of India, east coast of southern India. It is the List of states and union territories of India by area, seventh-largest state and th ...
, India
***Haplogroup M41c – found in Andhra Pradesh, India
*Haplogroup M2
– found in Melanesia
Melanesia (, ) is a subregion of Oceania in the southwestern Pacific Ocean. It extends from New Guinea in the west to the Fiji Islands in the east, and includes the Arafura Sea.
The region includes the four independent countries of Fiji, Vanu ...
*Haplogroup M2
– found in Melanesia and in a single Han individual from China
*Haplogroup M29'Q
**Haplogroup M2
– found in Melanesia
** Haplogroup Q (mtDNA), Haplogroup Qbr>
– found in Melanesia and Australia (Aboriginal peoples)
*Haplogroup M3
– found among the Onge people, Onge, in the Andaman Islands
**Haplogroup M31a
***Haplogroup M31a1
****Haplogroup M31a1a – Andaman Islands
****Haplogroup M31a1b – Andaman Islands
***Haplogroup M31a2
****Haplogroup M31a2a – northern India (Munda, ''etc.''), Myanmar
****Haplogroup M31a2b – India (Paudibhuiya)
**Haplogroup M31b'c
***Haplogroup M31b – northern India, Nepal, Myanmar
***Haplogroup M31c – Nepal (Tharu)
*Haplogroup M32'56 – Thailand
**Haplogroup M32
***Haplogroup M32a �
– found in the Andaman Islands
***Haplogroup M32c – Madagascar, Saudi Arabia, Thailand/Laos, USA
**Haplogroup M56 – India ( Korku)
*Haplogroup M33 – India
**Haplogroup M33a – found in Nepal (Tharu), India
***Haplogroup M33a1
****Haplogroup M33a1a – Lepcha, Tharu
****Haplogroup M33a1b – Dongri Bhil, Gujarat
***Haplogroup M33a2'3
****Haplogroup M33a2 – India (Katkari, ''etc.''), Egypt ( Siwi people, Siwa)
*****Haplogroup M33a2a – India, Iraq (Marsh Arab), Saudi Arabia
****Haplogroup M33a3 – found among Hindus in New Delhi, India
*****Haplogroup M33a3a – Myanmar, Toto
*****Haplogroup M33a3b – found in Thailand and in Tajiks in Dushanbe, Tajikistan
**Haplogroup M33b'c (M33-T16362C, M33-a) – northern India
***Haplogroup M33b – Nepal (Tharu), India
****Haplogroup M33b1 – Sonowal Kachari, Dai ( Jianshui)
****Haplogroup M33b2 – India, Nepal (Kathmandu)
***Haplogroup M33c – Ashkenazi Jews
Ashkenazi Jews ( ; also known as Ashkenazic Jews or Ashkenazim) form a distinct subgroup of the Jewish diaspora, that emerged in the Holy Roman Empire around the end of the first millennium CE. They traditionally speak Yiddish, a language ...
(Lithuania
Lithuania, officially the Republic of Lithuania, is a country in the Baltic region of Europe. It is one of three Baltic states and lies on the eastern shore of the Baltic Sea, bordered by Latvia to the north, Belarus to the east and south, P ...
, Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
, Belarus
Belarus, officially the Republic of Belarus, is a landlocked country in Eastern Europe. It is bordered by Russia to the east and northeast, Ukraine to the south, Poland to the west, and Lithuania and Latvia to the northwest. Belarus spans an a ...
, Ukraine
Ukraine is a country in Eastern Europe. It is the List of European countries by area, second-largest country in Europe after Russia, which Russia–Ukraine border, borders it to the east and northeast. Ukraine also borders Belarus to the nor ...
, Romania
Romania is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern and Southeast Europe. It borders Ukraine to the north and east, Hungary to the west, Serbia to the southwest, Bulgaria to the south, Moldova to ...
, Hungary
Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...
, Austria
Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...
, Latvia
Latvia, officially the Republic of Latvia, is a country in the Baltic region of Northern Europe. It is one of the three Baltic states, along with Estonia to the north and Lithuania to the south. It borders Russia to the east and Belarus to t ...
, Poland
Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...
), Han Chinese
The Han Chinese, alternatively the Han people, are an East Asian people, East Asian ethnic group native to Greater China. With a global population of over 1.4 billion, the Han Chinese are the list of contemporary ethnic groups, world's la ...
(Guangdong
) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
, Shaanxi
Shaanxi is a Provinces of China, province in north Northwestern China. It borders the province-level divisions of Inner Mongolia to the north; Shanxi and Henan to the east; Hubei, Chongqing, and Sichuan to the south; and Gansu and Ningxia to t ...
, Sichuan
Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...
, Hunan
Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
, Jiangsu
Jiangsu is a coastal Provinces of the People's Republic of China, province in East China. It is one of the leading provinces in finance, education, technology, and tourism, with its capital in Nanjing. Jiangsu is the List of Chinese administra ...
, Jilin
)
, image_skyline = Changbaishan Tianchi from western rim.jpg
, image_alt =
, image_caption = View of Heaven Lake
, image_map = Jilin in China (+all claims hatched).svg
, mapsize = 275px
, map_al ...
), Yao ( Jianghua, Guangxi
Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...
, Guangdong
) means "wide" or "vast", and has been associated with the region since the creation of Guang Prefecture in AD 226. The name "''Guang''" ultimately came from Guangxin ( zh, labels=no, first=t, t= , s=广信), an outpost established in Han dynasty ...
), Miao (Hunan
Hunan is an inland Provinces of China, province in Central China. Located in the middle reaches of the Yangtze watershed, it borders the Administrative divisions of China, province-level divisions of Hubei to the north, Jiangxi to the east, Gu ...
), Tibetans (Yunnan
Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...
), Zhuang (Guangxi
Guangxi,; officially the Guangxi Zhuang Autonomous Region, is an Autonomous regions of China, autonomous region of the China, People's Republic of China, located in South China and bordering Vietnam (Hà Giang Province, Hà Giang, Cao Bằn ...
), speakers of Kam-Tai languages (Guizhou
)
, image_skyline =
, image_caption =
, image_map = Guizhou in China (+all claims hatched).svg
, mapsize = 275px
, map_alt = Map showing the location of Guizhou Province
, map_caption = Map s ...
), Thai (Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
), Vietnamese
**Haplogroup M33d – India (Malpaharia, ''etc.'')
*Haplogroup M34'57
**Haplogroup M3
– small clade found in South Asia
***Haplogroup M34a – found in Karnataka
Karnataka ( ) is a States and union territories of India, state in the southwestern region of India. It was Unification of Karnataka, formed as Mysore State on 1 November 1956, with the passage of the States Reorganisation Act, 1956, States Re ...
, India
****Haplogroup M34a1 – India
*****Haplogroup M34a1a – India, Myanmar
****Haplogroup M34a2 – India (Pauri Bhuiya, Munda)
***Haplogroup M34b – India (Nihal, ''etc.'')
**Haplogroup M57 – India (Kathakur)
***Haplogroup M57a – India (Katkari, Nihal), England, Ireland
***Haplogroup M57b – India (Kathakur)
****Haplogroup M57b1 – India (Dongri Bhil)
*Haplogroup M3
– Nepal (Tharu)
**Haplogroup M35a – Tajiks in Xinjiang, Sarikoli, Armenia, Mesolithic Sri Lanka
Sri Lanka, officially the Democratic Socialist Republic of Sri Lanka, also known historically as Ceylon, is an island country in South Asia. It lies in the Indian Ocean, southwest of the Bay of Bengal, separated from the Indian subcontinent, ...
***Haplogroup M35a1 – India (Andh, Dongri Bhil), Mauritius
****Haplogroup M35a1a – India (Betta Kurumba, Mullukurunan)
***Haplogroup M35a2 – India (Andh, ''etc.'')
**Haplogroup M35b – found in Karnataka, Ladakh, Nepal (Tharu), Myanmar, Thailand, Slovakia
***Haplogroup M35b1'2'3 – India, Germany, USA
****Haplogroup M35b1 – India (Madia)
****Haplogroup M35b2 – India (Kathodi, Munda), Russia
****Haplogroup M35b3 – India (Ladakh)
***Haplogroup M35b4 – India ( Toto), Nepal (Kathmandu)
**Haplogroup M35c – India (Kathodi, Andh)
*Haplogroup M39'70
**Haplogroup M3
– found in South Asia
**Haplogroup M70 - Nepal ( Sherpas, ''etc.''), Tibet, Vietnam ( La Hủ)
*Haplogroup M4
– found in South Asia
**Haplogroup M40a – Yemen
***Haplogroup M40a1
****Haplogroup M40a1a
****Haplogroup M40a1b
*Haplogroup M42'74
**Haplogroup M4
– Sri Lanka
***Haplogroup M42a – Australian Aboriginal peoples
***Haplogroup M42b
****Haplogroup M42b1
****Haplogroup M42b2
**Haplogroup M74 - Cambodia ( Kampong Thom Province, Kampong Thom, Brao), Thailand ( Lao Isan, Central Thai), Vietnam ( Thái)
***Haplogroup M74a - Thailand ( Hmong, Iu Mien, Lao Isan from Nakhon Ratchasima Province), Vietnam ( Cờ Lao, Nùng, H'Mông, Mang), China, Taiwan
***Haplogroup M74b
****Haplogroup M74b1
****Haplogroup M74b2
*Haplogroup M4
– rare clade found in Saudi Arabia
*Haplogroup M49 – found among ancient specimens in the Euphrates valley
*Haplogroup M55'77
**Haplogroup M55 - Myanmar, Thailand, Malay
**Haplogroup M77 - Indonesia
*Haplogroup M62'68
**Haplogroup M62
**Haplogroup M68
***Haplogroup M68a1
****Haplogroup M68a1a - Cambodia, Vietnam
****Haplogroup M68a1b
***Haplogroup M68a2
****Haplogroup M68a2a- Cambodia
****Haplogroup M68a2b
****Haplogroup M68a2c
*Haplogroup M71 - India
**Haplogroup M71a'b (M71-C151T) - India, Myanmar, Cambodia ( Mel-Khaonh language, Mel), Laos (Lao in Vientiane), Thailand (Lawa, Karen, Shan, Blang, Phuan, Lao Isan, Khon Mueang), Vietnam ( Ede). Also found in approximately 10,500 year-old human remains ("Longlin") from Longlin Cave near De'e, Longlin County, Guangxi Province, China.
***Haplogroup M71a - Thailand ( Thai, Tai Yuan, Khon Mueang), China (Fujian
Fujian is a provinces of China, province in East China, southeastern China. Fujian is bordered by Zhejiang to the north, Jiangxi to the west, Guangdong to the south, and the Taiwan Strait to the east. Its capital is Fuzhou and its largest prefe ...
)
****Haplogroup M71a1 - China ( Han from Xiamen
Xiamen,), also known as Amoy ( ; from the Zhangzhou Hokkien pronunciation, zh, c=, s=, t=, p=, poj=Ē͘-mûi, historically romanized as Amoy, is a sub-provincial city in southeastern Fujian, People's Republic of China, beside the Taiwan Stra ...
and Lanzhou, Naxi)
*****Haplogroup M71a1a - China ( Bouyei from Pingtang, ''etc.''), Vietnam ( Mang, Kinh), Thailand ( Phuan from Central Thailand)
****Haplogroup M71a2 - India, Myanmar, Thailand ( Thai from Central Thailand and Eastern Thailand, Tai Yuan and Tai Khün from Northern Thailand, Blang), Indonesia, Philippines
The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
***Haplogroup M71b - Thailand ( Thai from Western Thailand, Khon Mueang from Chiang Mai Province, Tai Yuan, Tai Khün), China ( Bouyei from Pingtang, Han from Dongguan
Dongguan,; pinyin: alternately romanized via Cantonese as Tungkun, is a prefecture-level city in central Guangdong Province, China. An important industrial city in the Pearl River Delta, Dongguan borders the provincial capital of Guangzhou t ...
)
**Haplogroup M71c - Thailand ( Urak Lawoi, Moken, Thai from Central Thailand), Vietnam ( Cham from Bình Thuận, Kinh)
*Haplogroup M73'79
**Haplogroup M73
***Haplogroup M73a
****Haplogroup M73a1
*****Haplogroup M73a1a
*****Haplogroup M73a1b - Vietnam
****Haplogroup M73a2 - Papua New Guinea, East Timor
****Haplogroup M73a3 - Philippines (Aklan)
***Haplogroup M73b - Indonesia
****Haplogroup M73b1 - Vietnam, Cambodia, Indonesia
***Haplogroup M73c
**Haplogroup M79 - China
*Haplogroup M80'D
**Haplogroup M80 – found in Batak people of Palawan
Palawan (, ), officially the Province of Palawan (; ), is an archipelagic province of the Philippines that is located in the region of Mimaropa. It is the largest province in the country in terms of total area of . The capital and largest c ...
** Haplogroup D – found in Eastern Eurasia, Native Americans, Central Asia and occasionally also in West Asia and Europe.
Subclades
Tree
This phylogenetic tree of haplogroup M subclades is based on the paper by Mannis van Oven and Manfred Kayser ''Updated comprehensive phylogenetic tree of global human mitochondrial DNA variation'' and subsequent published research.
*M
**M1
***M1a
****M1a1
*****M1a1a
*****M1a1b
******M1a1b1
*****M1a1c
*****M1a1d
*****M1a1e
*****M1a1f
****M1a2
*****M1a2a
*****M1a2b
****M1a3
*****M1a3a
*****M1a3b
****M1a4
****M1a5
***M1b
****M1b1
*****M1b1a
****M1b2
*****M1b2a
**M2
***M2a
****M2a1
****M2a2
****M2a3
***M2b
****M2b1
****M2b2
**M3
***M3a
**M4"45
***M4
****M4a
****M4b
*****M4b1
***M18'38
****M18
****M38
*** M30
****M30a
****M30b
****M30c
*****M30c1
******M30c1a
*******M30c1a1
****M30d
***M37
****M37a
***M43
***M45
**M5
***M5a
****M5a1
*****M5a1a
*****M5a1b
****M5a2
*****M5a2a
**M6
**M7
***M7a
****M7a1
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*****M7e
**M8
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****M8a2
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*** CZ
**** C
**** Z
**M9
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*******M9a2
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*** E
**M10'42
***M10
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*****M10a2
***M42
****M42a
**M11
***M11a
***M11b
**M12'G
***M12
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*** G
**M13
***M13a
****M13a1
**M14
**M15
**M21
***M21a'b
****M21a
****M21b
***M21c'd
****M21c
****M21d
**M22
**M23
**M25
**M27
***M27a
***M27b
***M27c
**M28
***M28a
***M28b
**M29'Q
***M29
****M29a
****M29b
*** Q
**M31
***M31a
****M31a1
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*****M31b2
****M31c
**M32'56
***M32
****M32a
****M32c
***M56
**M33
***M33a
***M33b
***M33c
**M34
***M34a
**M35
***M35a
***M35b
**M36
***M36a
**M39
***M39a
**M40
***M40a
**M41
**M44'52
***M44
***M52
**M46
**M47'50
***M47
***M50
**M48
**M49
**M51
**M52’58
**M52
** D
Notable members
*Broadcast journalist Robert Siegel is a member of haplogroup M1a1b.
*Violinist Joshua Bell belongs to haplogroup M33c.
See also
* Genealogical DNA test
* Genetic genealogy
* Human mitochondrial genetics
*Population genetics
Population genetics is a subfield of genetics that deals with genetic differences within and among populations, and is a part of evolutionary biology. Studies in this branch of biology examine such phenomena as Adaptation (biology), adaptation, s ...
*Human mitochondrial DNA haplogroups
In human genetics, a human mitochondrial DNA haplogroup is a haplogroup defined by differences in Human mitochondrial genetics, human mitochondrial DNA. Haplogroups are used to represent the major branch points on the mitochondrial phylogenetic ...
* Red Deer Cave people
References
External links
*General
**Ian Logan'
Mitochondrial DNA Site
The India DNA
geographical project at Family Tree DNA
The China DNA
geographical project at Family Tree DNA
*Haplogroup M
**Mannis van Oven'
PhyloTree.org - mtDNA subtree M
**MITOMAP'
Haplogroup M
**FamilyTreeDNA's mtDNA Haplotree
Haplogroup M
from '' National Geographic''
Tree of M haplogroup as for 2006
Haplogroup M (mtDNA)
interest group on Facebook
Facebook is a social media and social networking service owned by the American technology conglomerate Meta Platforms, Meta. Created in 2004 by Mark Zuckerberg with four other Harvard College students and roommates, Eduardo Saverin, Andre ...
Another tree emphasizing the Andamanese and Nicobarese populations in comparison with other peoples with high M presence
** {{cite journal , last1=Thangaraj , first1=Kumarasamy , last2=Chaubey , first2=Gyaneshwer , last3=Singh , first3=Vijay Kumar , last4=Vanniarajan , first4=Ayyasamy , last5=Thanseem , first5=Ismail , last6=Reddy , first6=Alla G , last7=Singh , first7=Lalji , title=In situ origin of deep rooting lineages of mitochondrial Macrohaplogroup 'M' in India , journal=BMC Genomics , date=December 2006 , volume=7 , issue=1 , page=151 , doi=10.1186/1471-2164-7-151 , doi-access=free , pmid=16776823 , pmc=1534032
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