The grass carp (''Ctenopharyngodon idella'') is a

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

of large

herbivorous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...

freshwater fish Freshwater fish are those that spend some or all of their lives in fresh water, such as rivers and lakes, with a salinity of less than 1.05%. These environments differ from marine conditions in many ways, especially the difference in levels of s ...

in the

family Family (from la, familia) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its ...

Cyprinidae Cyprinidae is a family of freshwater fish commonly called the carp or minnow family. It includes the carps, the true minnows, and relatives like the barbs and barbels. Cyprinidae is the largest and most diverse fish family and the largest verte ...

, native to the

Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...

Far East The ''Far East'' was a European term to refer to the geographical regions that includes East and Southeast Asia as well as the Russian Far East to a lesser extent. South Asia is sometimes also included for economic and cultural reasons. The ter ...

, with a

native range Species distribution —or species dispersion — is the manner in which a biological taxon is spatially arranged. The geographic limits of a particular taxon's distribution is its range, often represented as shaded areas on a map. Patterns of ...

stretching from

northern Vietnam Northern Vietnam ( vi, Bắc Bộ) is one of three geographical regions within Vietnam. It consists of three administrative regions: the Northwest (Vùng Tây Bắc), the Northeast (Vùng Đông Bắc), and the Red River Delta (Đồng Bằng S ...

to the Amur River on the Sino-Russian border.Mandrak and Cudmore. 2004
''Biological Synopsis of Grass Carp (Ctenopharyngodon idella)''
. This Asian carp is the only species of the

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

''Ctenopharyngodon''. Grass carp are resident fish of large turbid

river A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of wate ...

s and associated

floodplain lake A lake is an area filled with water, localized in a basin, surrounded by land, and distinct from any river or other outlet that serves to feed or drain the lake. Lakes lie on land and are not part of the ocean, although, like the much larger ...

wetland A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (anoxic) processes prevailing, especially in the soils. The ...

s with a wide range of temperature tolerance, and spawn at temperatures of . It is cultivated as a

food fish Food is any substance consumed by an organism for nutritional support. Food is usually of plant, animal, or fungal origin, and contains essential nutrients, such as carbohydrates, fats, proteins, vitamins, or minerals. The substance is ing ...

China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's most populous country, with a population exceeding 1.4 billion, slightly ahead of India. China spans the equivalent of five time zones and ...

for centuries, but was introduced in

Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...

and the

United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie ...

for aquatic weed control, becoming the fish species with the largest reported farmed production globally, over five million tonnes per year.World aquaculture production of fish, crustaceans, mollusks, etc., by principal species in 2013
FAO Yearbook of Fisheries Statistics 2014

Appearance and anatomy

Grass carp have elongated, chubby, torpedo-shaped body forms. The terminal mouth is slightly oblique with non-fleshy, firm lips, and no

barbel Barbel may refer to: *Barbel (anatomy), a whisker-like organ near the mouth found in some fish (notably catfish, loaches and cyprinids) and turtles *Barbel (fish), a common name for certain species of fish **''Barbus barbus'', a species of cyprinid ...

s.''Grass carp''
in ''fishbase.org''. The complete lateral line contains 40 to 42 scales. Broad, ridged

pharyngeal teeth Pharyngeal teeth are teeth in the pharyngeal arch of the throat of cyprinids, suckers, and a number of other fish species otherwise lacking teeth.

Ecology

Grass carp inhabit lakes,

pond A pond is an area filled with water, either natural or artificial, that is smaller than a lake. Defining them to be less than in area, less than deep, and with less than 30% emergent vegetation helps in distinguishing their ecology from th ...

pool Pool may refer to: Water pool * Swimming pool, usually an artificial structure containing a large body of water intended for swimming * Reflecting pool, a shallow pool designed to reflect a structure and its surroundings * Tide pool, a rocky po ...

s and backwaters of large rivers, preferring large, slow-flowing or standing

water bodies A body of water or waterbody (often spelled water body) is any significant accumulation of water on the surface of Earth or another planet. The term most often refers to oceans, seas, and lakes, but it includes smaller pools of water such as p ...

with abundant vegetation. In the wild, grass carp spawn in fast-moving rivers, and their eggs, which are slightly heavier than water, develop while drifting downstream, kept in suspension by turbulence. Grass carp require long rivers for the survival of the eggs and very young fish, and the eggs are thought to die if they sink to the bottom. Adults grass carp feed primarily on

aquatic plant Aquatic plants are plants that have adapted to living in aquatic environments (saltwater or freshwater). They are also referred to as hydrophytes or macrophytes to distinguish them from algae and other microphytes. A macrophyte is a plant that ...

s, both higher aquatic plants and submerged terrestrial vegetation, but may also eat

detritus In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts commun ...

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs ...

s and other

invertebrate Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chordate ...

s. They eat up to three times their own body weight daily, and thrive in small lakes and backwaters that provide an abundant supply of vegetation. According to one study, grass carp live 5-9 years, with the oldest surviving 11 years. In

Silver Lake Silver is a chemical element with the Symbol (chemistry), symbol Ag (from the Latin ', derived from the Proto-Indo-European wikt:Reconstruction:Proto-Indo-European/h₂erǵ-, ''h₂erǵ'': "shiny" or "white") and atomic number 47. A soft, whi ...

Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...

, a thriving population of grass carp is passing the 15-year mark.

Introduced species

Grass carp have been introduced to many countries around the world. In the Northern Hemisphere, countries and territories of introduction include

Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...

, the

Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...

Malaysia Malaysia ( ; ) is a country in Southeast Asia. The federation, federal constitutional monarchy consists of States and federal territories of Malaysia, thirteen states and three federal territories, separated by the South China Sea into two r ...

India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...

Pakistan Pakistan ( ur, ), officially the Islamic Republic of Pakistan ( ur, , label=none), is a country in South Asia. It is the world's List of countries and dependencies by population, fifth-most populous country, with a population of almost 24 ...

Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...

Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...

, the

Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...

Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic country located on ...

Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Danish Realm, Kingdom of Denmark ...

, the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and North ...

France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...

Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...

, the

Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...

Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...

Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous ...

, the

Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...

Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the s ...

Romania Romania ( ; ro, România ) is a country located at the crossroads of Central Europe, Central, Eastern Europe, Eastern, and Southeast Europe, Southeastern Europe. It borders Bulgaria to the south, Ukraine to the north, Hungary to the west, S ...

Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = "Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capit ...

Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, an ...

Serbia Serbia (, ; Serbian language, Serbian: , , ), officially the Republic of Serbia (Serbian language, Serbian: , , ), is a landlocked country in Southeast Europe, Southeastern and Central Europe, situated at the crossroads of the Pannonian Bas ...

Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = M ...

Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and H ...

and Macedonia. In the Southern Hemisphere, they have been introduced to

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...

Australia Australia, officially the Commonwealth of Australia, is a Sovereign state, sovereign country comprising the mainland of the Australia (continent), Australian continent, the island of Tasmania, and numerous List of islands of Australia, sma ...

New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island count ...

Fiji Fiji ( , ,; fj, Viti, ; Fiji Hindi: फ़िजी, ''Fijī''), officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists ...

and

South Africa South Africa, officially the Republic of South Africa (RSA), is the southernmost country in Africa. It is bounded to the south by of coastline that stretch along the South Atlantic and Indian Oceans; to the north by the neighbouring countri ...

. Grass carp are known to have spawned and established self-reproducing populations in only six of the many larger Northern Hemisphere rivers into which they have been stocked. Their failure to establish populations in other rivers suggests they have quite specific reproductive requirements. In the United States, the species was first imported in 1963 from

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at the junction of the East and South China Seas in the northwestern Pacific Ocean, with the People's Republic of China (PRC) to the northwest, Japan to the nort ...

and Malaysia to aquaculture facilities in

Alabama (We dare defend our rights) , anthem = "Alabama (state song), Alabama" , image_map = Alabama in United States.svg , seat = Montgomery, Alabama, Montgomery , LargestCity = Huntsville, Alabama, Huntsville , LargestCounty = Baldwin County, Al ...

and

Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the Osage ...

. The first release is believed to have been an accidental escape in 1966 from the

U.S. Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...

's Fish Farming Experimental Station in

Stuttgart, Arkansas Stuttgart is a city in and the county seat of the northern district of Arkansas County, Arkansas, United States. Established by German settlers, it was named for its larger German counterpart. Known as the "Rice and Duck Capital of the World", t ...

, followed by planned introductions beginning in 1969. Subsequently, authorized, illegal and accidental introductions have been widespread; by the 1970s, the species had been introduced to 40 states, and it has since been reported in 45 of the country's 50 states. In 2013, it was determined to be reproducing in the

Great Lakes Basin The Great Lakes Basin consists of the Great Lakes and the surrounding lands of the states of Illinois, Indiana, Michigan, Minnesota, New York, Ohio, Pennsylvania, and Wisconsin in the United States, and the province of Ontario in Canada, whose dir ...

. It is still stocked in many states as an effective

biocontrol Biological control or biocontrol is a method of controlling pests, such as insects, mites, weeds, and plant diseases, using other organisms. It relies on predation, parasitism, herbivory, or other natural mechanisms, but typically also invo ...

for undesirable aquatic vegetation, many species of which are themselves introduced.

Use as weed control

Grass carp were introduced into New Zealand in 1966 to control the growth of aquatic plants. Unlike the other introduced fish brought to New Zealand, the potential value and impact of grass carp was investigated in secure facilities prior to their use in field trials. They are now approved by the New Zealand government for aquatic weed control, although each instance requires specific authorization."Grass carp for weed control"
DOC In the Netherlands, the species was also introduced in 1973 to control over-abundant aquatic weeds. The release was controlled and regulated by the Dutch Ministry of Agriculture, Nature, and Food Quality. In both of these countries, control is made easier because grass carp are very unlikely to naturally reproduce because of their very specific breeding requirements, but elsewhere, control is obtained by the use of sterile,

triploid Polyploidy is a condition in which the cells of an organism have more than one pair of ( homologous) chromosomes. Most species whose cells have nuclei ( eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contain ...

fish.

Fishing for grass carp

Grass carp grow large and are strong fighters when hooked on a line, but because of their vegetarian habits and their wariness, they can be difficult to catch via

angling Angling is a fishing technique that uses a fish hook or "angle" (from Old English ''angol'') attached to a fishing line to tether individual fish in the mouth. The fishing line is usually manipulated via a fishing rod, although rodless techniqu ...

. The

IGFA The International Game Fish Association (''IGFA'') is the leading authority on angling pursuits and the keeper of the most current World Record fishing catches by fish categories. Fishermen who are sport fishers are careful to follow their string ...

World record for a grass carp caught on line and hook is , caught in

Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedon ...

in 2009."Grass carp"

Online. Retrieved 22 June 2016. The fish are also popular

sport fish Game fish, sport fish or quarry refer to popular fish pursued by recreational anglers, and can be freshwater or saltwater fish. Game fish can be eaten after being caught, or released after capture. Some game fish are also targeted commercial ...

in areas where

bowfishing Bowfishing is a method of hunting fish that uses specialized archery equipment to lethally shoot and retrieve the animal. Fish are shot with a barbed arrow that is attached with a special line to a reel mounted on a bow or crossbow. Historically, ...

is legal. Where grass carp populations are maintained through

stocking Stockings (also known as hose, especially in a historical context) are close-fitting, variously elastic garments covering the leg from the foot up to the knee or possibly part or all of the thigh. Stockings vary in color, design, and transparen ...

as a biocontrol for noxious weeds, fishermen are typically asked to return any caught to the water alive and unharmed.

References

External links

Species Profile - Grass Carp (''Ctenopharyngodon idella'')
National Invasive Species Information Center,

United States National Agricultural Library The United States National Agricultural Library (NAL) is one of the world's largest agricultural research libraries, and serves as a national library of the United States and as the library of the United States Department of Agriculture. Located ...

. Lists general information and resources for Grass Carp. * * {{taxonbar, from=Q76098

grass carp The grass carp (''Ctenopharyngodon idella'') is a species of large herbivorous freshwater fish in the family Cyprinidae, native to the Pacific Far East, with a native range stretching from northern Vietnam to the Amur River on the Sino-Russian ...

Commercial fish Freshwater fish of China Fish of East Asia Herbivorous vertebrates Taxa named by Achille Valenciennes

Squaliobarbinae Chinese seafood dishes