Grace's Warbler
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Grace's warbler (''Setophaga graciae'') is a small
New World warbler The New World warblers or wood-warblers are a group of small, often colorful, passerine birds that make up the family Parulidae and are restricted to the New World. The family contains 120 species. They are not closely related to Old World warb ...
that specializes in
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
woods.


Taxonomy

Grace's warbler was discovered by
Elliott Coues Elliott Ladd Coues (; September 9, 1842 – December 25, 1899) was an American army surgeon, historian, ornithologist, and author. He led surveys of the Arizona Territory, and later as secretary of the United States Geological and Geographi ...
in the
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in great-circle distance, straight-line distance from the northernmost part of Western Can ...
in 1864. He requested that the new species be named after his 18-year-old sister, Grace Darling Coues, and his request was honored when
Spencer Fullerton Baird Spencer Fullerton Baird (; February 3, 1823 – August 19, 1887) was an American naturalist, ornithologist, ichthyologist, Herpetology, herpetologist, and museum curator. Baird was the first curator to be named at the Smithsonian Institution. He ...
described the species scientifically in 1865.


Description

Grace's warbler is a small bird, 11–13 cm long. It is mostly gray on top, with broken black streaks across the back and flanks and two white bars on the wings. The throat and breast are a vibrant yellow, and the rest of the underparts are white. It has a yellow half-eye ring under the eye and a long yellow "
eyebrow An eyebrow is an area of short hairs above each eye that follows the shape of the lower margin of the Supraorbital ridge, brow ridges of some mammals. In humans, eyebrows serve two main functions: first, human communication, communication thro ...
" stripe that starts at the base of the beak and runs above the eye, fading to white after it passes the eye.


Distribution and habitat

Grace's warblers breed from the south-western United States to Nicaragua and winter in their breeding range from north-eastern
Sinaloa Sinaloa (), officially the (), is one of the 31 states which, along with Mexico City, compose the Federal Entities of Mexico. It is divided into 18 municipalities, and its capital city is Culiacán Rosales. It is located in northwest Mexic ...
and north-western
Durango Durango, officially the Free and Sovereign State of Durango, is one of the 31 states which make up the Political divisions of Mexico, 32 Federal Entities of Mexico, situated in the northwest portion of the country. With a population of 1,832,650 ...
south. They are almost always found in pine forests, even in
migration Migration, migratory, or migrate may refer to: Human migration * Human migration, physical movement by humans from one region to another ** International migration, when peoples cross state boundaries and stay in the host state for some minimum le ...
, though at the northern end of their range (in southern
Nevada Nevada ( ; ) is a landlocked state in the Western United States. It borders Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the seventh-most extensive, th ...
and
Utah Utah is a landlocked state in the Mountain states, Mountain West subregion of the Western United States. It is one of the Four Corners states, sharing a border with Arizona, Colorado, and New Mexico. It also borders Wyoming to the northea ...
and southwestern
Colorado Colorado is a U.S. state, state in the Western United States. It is one of the Mountain states, sharing the Four Corners region with Arizona, New Mexico, and Utah. It is also bordered by Wyoming to the north, Nebraska to the northeast, Kansas ...
) they occur in
spruce A spruce is a tree of the genus ''Picea'' ( ), a genus of about 40 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal ecosystem, boreal (taiga) regions of the Northern hemisphere. ''Picea'' ...
-
fir Firs are evergreen coniferous trees belonging to the genus ''Abies'' () in the family Pinaceae. There are approximately 48–65 extant species, found on mountains throughout much of North and Central America, Eurasia, and North Africa. The genu ...
forest. They prefer open, "parklike" woods, and in some areas such as northern
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
they prefer an understory of Gambel's oak. In the United States and northern Mexico they typically occur at altitudes of ); farther south they may occur as low as 250 m. They are common in some localities, and in areas of the Mosquitia of Nicaragua where the only tree is the Caribbean pine, Grace's warblers may be the only bird seen.Stacier, Cynthia A.; Guzy, Michael J. 2002
Grace's Warbler (''Setophaga graciae'')
''The Birds of North America Online'' (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. Accessed 2012-03-1
Grace's Warbler is represented by four subspecies, including the nominate. ''Setophaga graciae'' (S. F. Baird, 1865). Southern Nevada, southern Utah, southwestern Colorado, northern New Mexico, and western Texas south through eastern Sonora and western Chihuahua. ''Setophaga graciae yaegeri'' (A. R. Phillips & Webster, 1961). West-central Mexico from southern Sinaloa, Durango, and Zacatecas south to Jalisco and Colima. ''Setophaga graciae remota'' (Griscom, 1935). Pacific coast from Michoacan through Guerrero, Oaxaca, and Chiapas into Guatemala, El Salvador, Honduras, and northern Nicaragua. ''Setophaga graciae decora'' (Ridgway, 1873). Belize.


Behaviour and ecology


Breeding

The nesting habits of Grace's warblers are largely unknown, as nests are very rarely found. The nest is a compact cup of plant fibers, the inside lined with hair and feathers, placed high above ground on a tree branch, usually
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
. The female lays 3 to 5 white or cream-colored eggs, speckled with brown, and ringed at the larger end.


Feeding

Like the vast majority of warblers, Grace's warbler eats only
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
s and
spider Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and ran ...
s, as far as is known from limited studies. It forages mostly high in pine trees (86% of its foraging time in a study in Arizona was in pines, but it foraged in oaks more in Mexico where
olive warbler The olive warbler (''Peucedramus taeniatus'') is a small passerine bird. It is the only member of the genus ''Peucedramus'' and the family Peucedramidae. This species breeds from southern Arizona and New Mexico, USA, south through Mexico to Nica ...
s also occur). It catches its food from the bases of needle clusters and the upper bark of branches no thicker than its body. It will often hover to inspect
pine cones A conifer cone, or in formal botanical usage a strobilus, : strobili, is a seed-bearing organ on gymnosperm plants, especially in conifers and cycads. They are usually woody and variously conic, cylindrical, ovoid, to globular, and have scales ...
for insect larvae. In Arizona it seldom flies to catch insects, but in Nicaragua it often does.


References


External links


Grace's warbler photo gallery
VIRE
Photo-High Res
{{Taxonbar, from=Q27075938 Grace's warbler Native birds of the Southwestern United States Birds of Central America Birds of Mexico Fauna of the Chihuahuan Desert Grace's warbler Grace's warbler Birds of the Sierra Madre Occidental