Geastrum Flexuosum
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''Geastrum'' (
orthographical variant In biology, within the science of Nomenclature, scientific nomenclature, i.e. the naming of organisms, an orthographical variant (abbreviated orth. var.) in botany or an orthographic error in zoology, is a spelling mistake, typing mistake or writi ...
''Geaster'') is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of puffball-like
mushroom A mushroom or toadstool is the fleshy, spore-bearing Sporocarp (fungi), fruiting body of a fungus, typically produced above ground on soil or another food source. ''Toadstool'' generally refers to a poisonous mushroom. The standard for the n ...
s in the family
Geastraceae Geastrales is an order of gasterocarpic basidiomycetes (fungi) that are related to Gomphales. The order contains the families Geastraceae, which includes the "earthstars" formerly placed in Lycoperdales or Phallales, and also Schenellaceae, Sc ...
. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers to the behavior of the outer peridium. At maturity, the outer layer of the fruiting body splits into segments which turn outward creating a star-like pattern on the ground. The inner peridium is called a spore sac. In some species, the outer peridium splits from a middle layer, causing the spore sac to arch off the ground. If the outer peridium opens when wet and closes when dry, it is described as hygroscopic. In some species, the inner peridium is borne on a stalk or pedicel. The columella is a column-like clump of sterile tissue found inside the inner peridium. The network of fertile tissue inside the inner peridium, the
capillitium Capillitium (pl. capillitia) is a mass of sterile fibers within a fruit body interspersed among spores. It is found in Mycetozoa Mycetozoa is a polyphyletic grouping of slime molds. It was originally thought to be a monophyletic clade, but in ...
, arises from the columella. This is where
basidia A basidium (: basidia) is a microscopic spore-producing structure found on the hymenophore of reproductive bodies of basidiomycete fungi. The presence of basidia is one of the main characteristic features of the group. These bodies are also ...
and
basidiospores A basidiospore is a reproductive spore produced by basidiomycete fungi, a grouping that includes mushrooms, shelf fungi, rusts, and smuts. Basidiospores typically each contain one haploid nucleus that is the product of meiosis, and they are pr ...
are produced. The mouth in most species of "earth-stars" is quite prominent, often arising as a small cone at the apex of the inner peridium. It may be even or sulcate (grooved). They are generally not toxic but considered non-edible due to their fibrous texture in the mature stage at which they are generally found.


Species

Although the ''Dictionary of the Fungi'' (2008) estimated roughly 50 species in ''Geastrum'', a more recent estimate (2014) suggests that there may be up to 120 species. ''Geastrum coronatum'' Pers. has been proposed as the conserved type for the genus. Some similar species that are otherwise difficult to differentiate using classical morphological features (such as ''G. triplex'', ''G. saccatum'', and ''G. lageniforme'') can be identified using chemical spot tests that detect
phenoloxidase Polyphenol oxidase (PPO; also polyphenol oxidase i, chloroplastic), an enzyme involved in fruit browning, is a tetramer that contains four atoms of copper per molecule. PPO may accept monophenols and/or ''o''-diphenols as substrates. The ...
enzymatic activity, a differences in the crystal structure of
calcium oxalate Calcium oxalate (in archaic terminology, oxalate of lime) is a calcium salt of oxalic acid with the chemical formula or . It forms hydrates , where ''n'' varies from 1 to 3. Anhydrous and all hydrated forms are colorless or white. The monohydr ...
deposits or DNA sequences. Species include: * '' Geastrum aculeatum'' * '' Geastrum albonigrum'' * ''
Geastrum andrewsii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum argentatum'' * '' Geastrum argentinum'' * ''
Geastrum australe ''Geastrum australe'' is an inedible species of mushroom belonging to the genus ''Geastrum'', or earthstar fungi. It can be mistaken for ''Geastrum saccatum, G. saccatum'' or ''Geastrum simulans, G. simulans'', and is found in Melbourn ...
'' * ''
Geastrum austrominimum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum benitoi ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum britannicum ''Geastrum britannicum'' is a fungal species in the family Geastraceae. Its recommended English name is vaulted earthstar. Like other earthstars, the basidiocarps (fruit bodies) are initially globose. Their thick outer skin splits open at maturit ...
'' * ''
Geastrum berkeleyi ''Geastrum berkeleyi'', or Berkeley's earthstar, is an inedible species of mushroom belonging to the genus ''Geastrum'', or earthstar fungi. It can be distinguished from other ''Geastrum'' species by the flat bipyramidal shape of the calcium oxala ...
'' * ''
Geastrum bushnellii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum campestre ''Geastrum campestre'' is an inedible species of mushroom belonging to the genus ''Geastrum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as e ...
'' * ''
Geastrum clelandii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum congolense ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * ''
Geastrum corollinum ''Geastrum corollinum'' is an inedible species of mushroom belonging to the genus ''Geastrum'', or earthstar fungi. First described scientifically by German naturalist August Johann Georg Karl Batsch in 1792 as ''Lycoperdon corollinum'', it was t ...
'' * '' Geastrum coronatum'' * ''
Geastrum dissimile ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum drummondii'' * '' Geastrum dubowskii'' * '' Geastrum echinulatum'' * '' Geastrum elegans'' * '' Geastrum elliptice'' * '' Geastrum entomophilum'' * '' Geastrum episcopale'' * '' Geastrum fimbriatum'' * '' Geastrum flexuosum'' * '' Geastrum floriforme'' * '' Geastrum fornicatum'' * '' Geastrum fuscogleba'' * '' Geastrum hieronymi'' * '' Geastrum hirsutum'' * '' Geastrum huneckii'' * '' Geastrum hungaricum'' * '' Geastrum inpaense'' – Brazil * '' Geastrum jurei'' * '' Geastrum kotlabae'' * ''
Geastrum kuharii ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum lageniforme'' * '' Geastrum leptospermum'' * '' Geastrum lilloi'' * '' Geastrum litchiforme'' * '' Geastrum lloydianum'' * '' Geastrum magnosporum'' * '' Geastrum melanocephalum'' * '' Geastrum meridionale'' * '' Geastrum minimum'' * '' Geastrum mirabile'' * '' Geastrum morganii'' * '' Geastrum ovalisporum'' * '' Geastrum oxylobum'' * '' Geastrum papinuttii'' * '' Geastrum parvisporum'' * '' Geastrum parvistriatum'' * '' Geastrum pectinatum'' * '' Geastrum pleosporus'' * '' Geastrum pouzarii'' * '' Geastrum pseudolimbatum'' * '' Geastrum quadrifidum'' * '' Geastrum reticulatum'' * '' Geastrum rhizophorum'' * '' Geastrum rufescens'' * '' Geastrum rugulosum'' * '' Geastrum rusticum'' – Brazil * '' Geastrum saccatum'' * '' Geastrum schmidelii'' * '' Geastrum schweinitzii'' * '' Geastrum senoretiae'' * '' Geastrum setiferum'' * '' Geastrum smardae'' * '' Geastrum smithii'' * '' Geastrum stiptatum'' * ''
Geastrum striatum ''Geastrum'' (orthographical variant ''Geaster'') is a genus of puffball-like mushrooms in the family Geastraceae. Many species are commonly known as earthstars. The name, which comes from ''geo'' meaning ''earth'' and meaning ''star'', refers ...
'' * '' Geastrum subiculosum'' * '' Geastrum tichifer'' * ''
Geastrum triplex ''Geastrum triplex'' is a fungus found in the detritus and leaf litter of hardwood forests around the world. It is commonly known as the collared earthstar, the saucered earthstar, or the triple earthstar—and less commonly by the alternativ ...
'' * '' Geastrum velutinum'' * '' Geastrum verrucoramulosum'' * '' Geastrum welwitschii'' * '' Geastrum xerophilum'' * '' Geastrum xylogenum''


References


Sources


New Zealand Species

British Species

Czech Species and Photos
* *
Ponce de Leon, Patricio, A Revision of the Family Geastraceae, Field Museum of Natural History 1968
* * {{Taxonbar, from=Q687050 Agaricomycetes genera